Devonian sharks from Antarctica 
299 
specimens, and have more widely splayed main 
cusps and, relative to the holotype, they show a 
more curved, and distictiy shorter, mesial 
crenulated area when viewed in labial or lingual 
aspect. WAM 92.3.58 shows the presence of a small 
accessory cuspule on the external edge of the main 
cusp, a feature seen on nearly all specimens with 
that edge well-preserved (e.g., Figure 9J), although 
absent on 92.3.59 (Figure 8H, 9C). It is possible that 
these two teeth could represent a different, 
younger species of Aztecodus, although as only two 
specimens are known, and they show only slight 
differences from the other specimens, we prefer to 
place them in the same species until more material 
can substantiate or refute these morphological 
differences. 
Stratigraphic occurrence 
The type material comes from the Lashly Ranges, 
(section "LA"); other specimens are from Mt 
Ritchie, section A4 near units 61, 62; from Alligator 
Peak, locality 20 (top horizon) and from Fault Bluff, 
"fish hotel" site B, Cook Mountains. The teeth all 
come from above the thelodont biozones, with first 
appearance high in the section at "fish hotel" (at 
least 65 m from base of unit covered by scree, 
occurring with phyllolepids, Groetilandaspis 
antarcticus, Portalodus bradshawae gen. et sp. nov., 
and ranging right through to about 20 m from the 
top of the Aztec Siltstone at Alligator Ridge, site 20 
(possibly uppermost portalensis or karawaka 
biozones). 
Anareodus gen. nov. 
Anareodus statei sp. nov. 
Figures 8J-K, 10 
Etymology 
After ANARE (Australian National Antarctic 
Research Expeditions) who funded J. Long's field 
work, and for Brian Staite, survival leader on 
K221/A136 Expedition. 
Diagnosis 
A diplodont shark tooth with main cusps of very 
unequal size, separated by a mesial-marginal ridge 
sometimes bearing small crenulations, and a small 
accessory cusplet normally present on the mesial 
side of the main cusp. Main cusp quite flat and 
sigmoidally curved in lateral/mesial view. Base 
low, strongly concave, and rectangular in outline 
in basal view. 
Remarks 
This genus resembles Aztecodus harmsenae gen. et 
sp. nov. in sometimes showing crenulations on the 
central cutting ridge, in the presence of a small 
Figure 10 Anareodus statei gen. et sp. nov. All specimens, except D, shown in labial view; D, in lateral view. A, WAM 
90.2.38, from "Gorgon's Head" (Cook Mountains). B, WAM 94.2.13. C, WAM 94.2.9 (both from Fault Bluff, 
"fish hotel" B horizon). E, WAM 90.2.39 (Gorgon's Head). All bar scales are 1 mm. 
