Records of the Western Australian Museum 17: 309-314 (1995). 
The Hipposideros bicolor group (Chiroptera: Hipposideridae) from 
Sumbawa Island, Nusa Tenggara, Indonesia 
D.J. Kitchener 1 and Maharadatunkamsi 2 
1 Western Australian Museum, Francis Street, Perth, Western Australia 6000 
2 Museum Zoologicum Bogoriense, LIPI, Jin. Ir. H. Juanda 9, Bogor, Indonesia 16122 
Abstract - Two unique specimens of Hipposideros from West Sumbawa are 
described. 
INTRODUCTION 
Surveys of islands in the Greater and Lesser 
Sundas and the Maluku Tenggara region of 
Indonesia by staff from the Western Australian 
Museum and Museum Zoologicum Bogoriense, 
between 1987 and 1993, has resulted in the 
recognition and/or redefinition of a number of 
mammalian taxa (Kitchener et al. 1993 a,b). In 1988, 
two specimens of a unique Hipposideros bat were 
collected in West Sumbawa. 
The morphology of the above two West 
Sumbawa specimens are herein described. They 
belong in the H. bicolor group of Hill (1963). This 
group is characterised by: small to moderate body; 
large, broadly rounded ears, normally with an 
internal fold or thickening of the membrane of the 
ear at the antitragal lobe; elongate, narrow skull; 
moderately inflated braincase; narrow rostrum; 
unspecialised auditory region; upper incisors weak 
with outer lobe obsolescent or obsolete; and outer 
lower incisor crown dimensions subequal to those 
of the inner lower incisor. 
METHOD 
Measurements were recorded with vernier 
calipers. For skull, dentary, dental and baculum, 
measurements to 0.01 mm; for externals to 0.1 mm. 
Terminology used in the description of characters 
follows Hill (1963) and Smith (1984). Measurement 
points, where not explicit, are detailed in Kitchener 
and Maryanto (1993). Pelage colour description, 
when following the terminology of Smithe (1975), 
are capitalised. 
SYSTEMATICS 
Hipposideros sp. indet. 
Material examined 
Museum Zoologicum Bogoriense No. MZB 
15905, adult male carcase weighing 9.5 g; fixed in 
10% formalin and preserved in 75% ethanol; skull 
and mandible separate; liver stored in ultrafreeze 
at the Western Australian Museum (WAM); 
collected by D.J. Kitchener, R.A. How and 
Maharadatunkamsi, on 26 May 1988. WAM M 
31489; adult male weighing 9.8 gm; carcase and 
liver preserved as for the holotype; skull removed 
and lost. 
Locality 
Collected from a small cave near the main 
limestone "cathedral" cavern of Gua (=cave) Batu 
Tering, 3 km S Desa Batu Tering, West Sumbawa 
(c. 8°48'S, 117°22’E), at an altitude of ca. 200 m. A 
stream passes within a few metres of this small 
cave. The surrounding vegetation is dense gallery 
rainforest. 
Characteristics 
A member of the H. bicolor group (sensu Hill 1963) 
but separated from all other members of that 
group by the following combination of characters: 
moderately large size with forearm length 47.9- 
48.0, condylocanine length 16.7, and C l M 3 length 
5.7; lateral supplementary leaflets on face absent; 
anterior upper premolar (PM 2 ) present, only 
slightly extruded from toothrow but separates 
canine from PM 1 ; anterior lower premolar (PM 2 ) 
length subequal to second lower premolar (PM 4 ); 
PM, height three-quarters that of PM 4 ; anterior leaf 
without median emargination; posterior leaf with 
three well defined septa; intemarial septum slightly 
thickened; interorbital region constricted; glandular 
ridge on muzzle beneath margin of anterior leaf 
absent; anterior half of zygomata not particularly 
slender; superior projection of zygomata poorly 
developed; vomer posterior projection into 
mesopterygoid fossa blade-like, only very slightly 
thickened; and baculum moderately long (3.2), 
straight, narrow, bifurcated at distal tip with 
cranial proximal (projecting) margin of base evenly 
rounded and not bifurcated. 
