D.J. Kitchener, Maharadatunkamsi 
Figure 1 Location of Sumbawa Island and the locality of Hipposideros sp. indet. 
Skull (Figure 2) 
Moderately long, with greatest skull length 18.36 
and condylocanine length 16.72, and slender; 
zygomatic width 9.43, considerably greater than 
mastoid width 9.05; cranium comparatively broad 
8.81 and inflated; interorbital region much 
narrower than distance between anteorbital 
foramina 2.84 v. 5.15; sagittal crest low, reaches 
maximum height at cranial apex; rostral eminences 
slightly inflated, in lateral profile a little below 
junction of sagittal and supraorbital crests; in 
lateral profile, rostrum anterior edge slopes slightly 
downwards anteriorly and curves smoothly to 
maxilla; zygomata robust, superior jugal projection 
low; anteorbital foramen oval, closed by 
moderately wide bar of bone; premaxillae broad 
and closely oppose lingual margin of upper canine; 
premaxillae junction with a V shape; palate 
posterior margin a broad V shape without a 
median posterior spicule; mesopterygoid fossa 
narrow 1.74, projecting vomer blade-like, only very 
slightly thickened; sphenoidal bridge wide, viewed 
ventrally almost conceals sphenorbital sinus; 
sphenoidal depression, shallow, oval and wider 
than mesopterygoid fossa 3.02 v. 1.74; cochlea (the 
cochlea part of the petrosal) size moderate, their 
breadth considerably greater than their distance 
apart 2.64 v. 1.63; cochlea inflated, reaches depth 
level with tympanic bulla; tympanic bulla short, 
subequal in length to cochlea breadth, terminate c. 
0.2 posterior to glenoid fossa margin. 
Dentition 
Upper incisors small, their tips markedly 
convergent, outer lobe obsolescent; anterior upper 
premolar (PM 2 ) small, basal surface area 
approximately that of upper incisor, slightly 
extruded from toothrow but such that C 1 is not in 
contact with PM 4 ; M 3 premetacrista half length of 
its preparacrista; M 2 and M 3 with pronounced 
hypocone; M 1 components of eocrista shorter than 
those of M 2 but lengths of these teeth subequal; 
outer lower incisor crown area about 15% larger 
than that of inner lower incisor; anterior lower 
premolar (PM 2 ) 80% length and three-quarters 
height of PM 4 . 
Externals (Figure 3) 
Ears of moderate length 18.7-19.3, rounded, 
anterior margins strongly convex, posterior margin 
with very slight concavity immediately behind 
apex; antitragus with distinct fold; noseleaf simple; 
anterior noseleaf narrower than posterior noseleaf 
ca. 5.1 v. 5.8; lateral supplementary facial leaflets 
absent; internarial septum slightly expanded 
laterally, more so anteriorly to a maximum breadth 
of 0.8, vertically it is raised in a semicircle to a 
height of c. 1.0; laterally separated from anterior 
edge of nares by deep grooves; narial lappets 
border external narial margin, raised c. 0.4 above 
surface of anterior noseleaf; intermediate leaf 
unspecialised; posterior leaf supported by three 
prominent septa. 
Fur long, mid dorsal c. 8.5, mid rump c. 12.0 
chest c. 7.5, external pinna furred to apex c. 9.0. 
dorsum predominantly Cinnamon colour of distal 
one-quarter of hairs of which basal part a dull 
White; face Tawny. Venter, including chin, pale 
Smoke Gray; patagia Olive Brown. 
Penis long 8.8, lightly furred; glans penis 
flattened in craniocaudal axis, distal tip evenly 
semicircular except for two small projecting lobes 
surrounding the urethral opening (Figure 4). 
Baculum moderately long 3.22, thin, straight 
