Australian Pionidae 
375 
disto-dorsal seta; tarsus with 4 disto-dorsal setae. 
Legs (Figures 38-40): <J tarsus III with shortened 
and thickened tarsus, and with unequal claws, 
posterior claw shortest (Figure 39); 8 metatarsus 
III with row of spatulate setae on posterio-distal 
margin; 8 patella and tibia IV thickened, tibia with 
6 stout forward-projecting thick setae on postero- 
ventral margin; 8 metatarsus IV with large, thick 
protuberance extending over tarsus, about one- 
third as long as tarsus, concave edge of metatarsus 
with several small spinules (Figure 40); legs II, III 
and IV with swimming setae arranged as follows: 
leg II: 8, tibia 2-4, metatarsus 2-3; 2, tibia 2-4, 
metatarsus 4; leg III: 8, patella 0-2, tibia 6-8, 
metatarsus 0; 2, patella 0-2, tibia 4-5, metatarsus 
5; leg IV: 8, patella 0-1, tibia 3-5; 2, patella 0-3, 
metatarsus 5-7, tarsus 6-8. 
Dimensions (pm), 8 (2): body 565-710/455-586 
(730-1064/518-604); capitulum length 147-173 
(179-195); chelicera length 200 (?); genital field 118- 
134/288-294 (159-243/248-346). Pedipalp: 
trochanter 38-43 (41-50), femur 89-103 (106-121), 
patella 44-56 (58-65), tibia 80-105 (111-122), tarsus 
60-71 (65-72). Leg I: trochanter 70-88 (77-83), 
femur 71-90 (82-96), patella 99-121 (128-135), tibia 
141-163 (179-189), metatarsus 147-180 (197-205), 
tarsus 192-230 (202-250). Leg III: metatarsus 160- 
200, tarsus 127-144. Leg IV: trochanter 138-160 
(128-149), femur 108-134 (110-125), patella 138-186 
(173-186), tibia 122-163 (224-232), metatarsus ISO- 
147 (250-261), tarsus 237-288 (237-275). 
Deutonymph 
Much as in adults except that vgx is absent; 
genital region with 2 pairs of acetabula. 
Dimensions (pm): body 448/352. 
Remarks 
Lundblad (1941 and several other papers) did not 
make valid holotype designations or restriction of 
type localities for many species, and there is some 
doubt as to the type status of the female specimen 
from Mt Macedon, Victoria which he had labelled 
as the allotype. As this locality was not mentioned 
in the original publication, I treat it as not part of 
the type series, which is restricted to the holotype 
from Kalgoorlie. 
Males of A. falcipes differ from those of A. 
poorginup by the shape of the fourth leg and the 
lack of sclerotization posterior to the genital field. 
Both males and females differ by the lack of ventral 
setiferous tubercles on the pedipalpal tibia. 
Acercella falcipes is a remarkably widespread 
species (Figure 52), and only slight differences in 
size could be detected between eastern and 
western populations. The distribution conforms 
quite well to a Bassian distribution, and tire species 
may eventually be found in suitable habitats in 
Tasmania. 
During August 1987, several pairs collected from 
North Lake, W.A., were transferred to watch- 
glasses in the laboratory where mating was 
observed. The male approached the female and, 
with their ventral surfaces in contact, he clasped 
the female's fourth leg between the tarsus and the 
distal extension of the tarsus of his fourth leg. He 
then collected the spermatophore from his 
gonopore with his cupped third tarsi, and 
transferred it into the female's gonopore. These 
constitute the first observations on mating in the 
genus. 
Acercella poorginup sp. nov. 
Figures 42-52 
Material Examined 
Holotype 
8, Poorginup Swamp, Western Australia, 
Australia, 34°33'S, 116°44'E, 4 September 1985, M.S. 
Harvey, T.J. Doeg (WAM 88/2943; SL). 
Paratypes 
Australia: Western Australia: Id, 1$, same data 
as holotype (WAM 88/2944-2945; SL); 18, 12, 
same data as holotype (NMV K896-897; SL). 
Diagnosis 
Pedipalpal tibia with 2 large ventral setiferous 
tubercles. Male metatarsus IV with dorsal blade¬ 
like expansion very thin. 
Description 
Adult 
Colour deep red. Glandularia: 5 pairs of 
dorsoglandularia, 4 pairs of lateroglandularia, and 
6 pairs of ventroglandularia present (Figures 42- 
45). Dorsum (Figures 42, 44): with 2 pairs of 
dorsalia, anterior pair larger than posterior pair. 
Venter (Figures 43, 45): apodemes of coxa I very 
short; suture lines between coxae I and II and coxae 
III and IV present; suture lines between coxae III 
and IV extending laterally (8) or antero-laterally 
(2). Genital region (Figures 46, 47): 3 pairs of 
acetabula; those of 8 unequal in size, the posterior 
pair larger than remaining pairs; those of 2 
virtually in a straight line. Chelicera not examined. 
Pedipalp (Figure 51): tibia with 2 large disto- 
ventral setiferous tubercles, and with disto-dorsal 
seta; tarsus with 4 disto-dorsal setae. Legs (Figures 
48-50): 8 tarsus III extremely shortened and 
thickened, and with unequal claws, posterior claw 
shortest (Figure 49); 8 tibia and metatarsus IV 
modified, tibia with fan-shaped arrangement of 
swimming hairs on ventral surface, and metatarsus 
with large dorsal extension curving over tarsus. 
