Australian Pionidae 
379 
1836), by subsequent designation of Koch 
(1842). 
Curvipes Koenike, 1891: 20. Replacement name for 
Nesaea Koch, 1836. Synonymised by Wolcott 
(1905). 
Piona (Dispersipiona) K. Viets, 1926: 194. Type 
species: Hydrachna davicomis Muller, 1776, by 
original designation. Synonymised by Cook 
(1960). 
Piona (Telrapiona) K. Viets, 1926: 194. Type species: 
Nesaea variabilis Koch, 1836, by original 
designation. Synonymised by Cook (1960). 
Piona (Carnepiom) Besseling, 1959: 22. Type species: 
Nesaea carnea Koch, 1836 by original 
designation. Synonymised by Cook (1974). 
Piona (Pusillopiona) Besseling, 1959: 22. Type 
species: Nesaea pusilla Neuman, 1875 by original 
designation. Synonymised by Cook (1974). 
Piona (Conglopiona) Besseling, 1959: 22. Type 
species: Nesaea conglobata Koch, 1836 by original 
designation. Synonymised by Cook (1974). 
Diagnosis 
Male tarsus III with dissimilar claws; tibia IV of 
male with concavity lined with small, blunt setae. 
Body without accessory sclerotization. Genital field 
with many pairs of acetabula. Excretory pore 
surrounded by sclerotized ring. 
Remarks 
Due to the large number of described Piona 
species, and a certain amount of variation within 
the genus, several authors have erected subgenera. 
The history and status of these names was 
summarized by Cook (1974) who noted that their 
use was hazardous due to gradations between 
species. Smith (1976) adopted a more reasonable 
approach, and erected a series of species-group 
categories. 
Piona cumberlandensis (Rainbow) 
Figures 52-54 
A tax cwnberlandensis Rainbow, 1906: 160, fig. 37 (in 
part). 
Unionicola cumberlandensis (Rainbow): Walter, 1928: 
59; K. Viets, 1956: 360. 
Unionocola [sic] cumberlandensis (Rainbow): K. Viets, 
1932: 364; 
Unionicola (Hexatax) cumberlandensis (Rainbow): 
Halik, 1941: 103. 
Piona uncatiformis Lundblad, 1941: 118. 
Synonymized by Cook (1986). 
Piona (Piona) uncatiformis Lundblad: Lundblad, 
1947: 69, figs 44a-i; K. Viets, 1956: 460. 
Piona cumberlandensis (Rainbow): Cook, 1986: 201, 
figs 1053-1058, 1061. 
Figures 53-54 Piona cumberlandensis (Rainbow), $: 53, dorsal aspect; 54, ventral aspect. Scale line = 100 pm. 
