396 
J.H. Bradbury, W.D. Williams 
dimorphism in both the possession of antennal 
calceoli and of notched gnathopodal palms. 
In this paper we accept Liagoceradocus as a valid 
genus and ascribe two new species to it: L. 
subthalassicus and L. branchialis. The presence of the 
genus in the Indian Ocean considerably extends its 
known area of distribution. 
Liagoceradocus subthalassicus sp. nov. 
Figures 1-3 
Material Examined 
Holotype 
9 3.5mm. Unique (WAM 258-95, collection 
BES546). 
Type locality 
Ledge Cave B-l, an anchialine cave on Barrow 
Island, Western Australia, Australia (20°48'S, 
115°20'E), collected by trapping, 28 July 1992, W.F. 
Humphreys and B. Vine. 
Diagnosis 
Pleonites naked except for 3 small dorsal setae on 
pleonite 4. Head. Rostrum weak, lateral cephalic 
lobes moderately projecting, antennal sinus small 
and distinct, eyes absent. First antenna elongate, 
longer than A2, ratio of peduncular articles 
23:21:11, accessory flagellum 2 articulate, no 
calceoli. Second antenna short, flagellum much 
shorter than peduncle, calceoli absent. Upper lip. 
Apical margin rounded, symmetrical. Mandible. 
Palp 3 articulate, ratio of articles 17:39:41, article 1 
not setose, article 2 bearing 4 strong medial setae, 
article 3 sub-falciform, almost linear, setate, 
approximately equal in length to article 2, setae = 
D9,E4. Accessory blades (rakers) = 9, without inter¬ 
raker plumose setae but 1 additional short naked 
seta proximally. No additional serrations beyond 
rakers. Lower lip bearing small vestigial inner lobes. 
Maxillae moderately setose medially. First maxilla 
inner plate triangular or semi-circular, linear 
medially, with 12 plumose medial setae, outer plate 
with 7 spines, palps asymmetric, both plates well 
covered in pubescence. Second maxilla inner plate 
with oblique facial row of long setae, baso-medial 
and outer-distal margins, and apical margin of the 
outer plate pubescent, both plates with moderate 
to strong apical setae. Maxilliped. Inner plate long, 
extending beyond M0.5 of the outer plate, laterally 
pubescent, with a distal row of 5 plumose setae, 3 
blunt naked tooth spines, 1 sub-distal medial spine 
on the left side only, and 1 sub-marginal sub-distal 
medial spine forming a short row with 2 long 
plumose setae at M0.5 and MOV. Outer plate broad 
with 5 naked and 2 plumose distal setae, 2 strong, 
short distal sub-marginal medial tooth spines, the 
disto-medial margin carved into 2 or 3 acuminate 
sinuosities merging basaliy with a medial row of 8 
slender spines which become sub-marginal distally 
leaving the sinuosities naked. Palp article 2 without 
lateral setae, article 3 with a single lateral seta at 
M0.9, and 2 pairs of latero-facial setae at M0.4 and 
MOV, articles 2-3 moderately setose medially, 
article 3 bearing a transverse band of pubescence 
at M0.5, and distal comb rows of setae toward the 
base of the dactyl, the apex is moderately 
produced, dactyl unguiform with 3 distal accessory 
spinules, the nail of moderate length, bearing a 
single dorsal spinule of equal length. Coxae. Cl-3 
longer than broad, C4 as broad as long, Cl-4 with 
few posterior spines. Cl not expanded below, C4 
not excavate posteriorly. Cnathopods subchelate, 1— 
2 diverse. Setae of articles 4 ordinary, not rastellate. 
First gnathopod small, feeble, carpus longer than 
propodus, not lobate, merus not lobate but 
posteriorly bulbous and pubescent, left palm 
slightly oblique, right palm transverse, both short, 
palmar spines not symmetrically bifid but with 
small sub-apical trigger like extensions. Spines at 
comer of palm = 2 stout bifid medial and lateral, 
and 1 elongate, slender, lateral. Few spines or setae 
along the palm, dactyls lacking recumbent inner 
tooth spines, nail present. Second gnathopod 
enlarged, 2x Gl, carpus equal to propodus, with 
long keel like pubescent posterior lobe, posterior 
setae sub-marginal proximally, to facial distally, 
hand ovate, palm oblique, moderately spinose and 
setulose, defining comer with 2 stout medial spines 
and 2 elongate lateral spine-setae, nail absent, free 
posterior margin bearing long 'hadziid' setae 
(Barnard and Barnard, 1983). Pereopods feeble, 
weak. P3-4 smaller, weaker than P5-7, posterior 
spine sets on article 6 of P3-A weak, uneven. P5—7 
moderately elongate, P7 longest, P5 shortest, article 
2 moderately expanded, ovate, slightly lobate on 
P6, lobate on P7. Dactyls of P3-7 bearing 2 or more 
distal accessory spinules as well as an outer basal 
penicillate setule. Gills. Coxae 2-6 each with a 
single flask shaped gill. Gill 6 not reduced. 
Oostegites. Coxae 2-5 each bearing a thin, poorly 
setose oostegite. 
Epimera. Posteroventral tooth of El-3 absent 
except for a tiny spine on E3. Posterior margins 
smooth and naked except for 1 or 2 small setae. 
E2-3 with single, small posterior ventro-facial 
spinules and 1 or 2 antero-facial spines. Pleopods 
with dissimilar numbers of setae and retinacula. 
Rami extending sub-equally and symmetrically, 
baso-medial setae not bifid. Urosome. With 3 
ventro-distal spines on urosomite 1 at the base of 
UL Uropods. U3 extended, magniramous, 
dispariramous, peduncle short, outer ramus 2 
articulate. Apico-lateral comer of peduncles of Ul- 
2 with 1 spine, the dorsal margins spinose, medial 
margin of peduncle of U1 with 1 apical spine, of 
