402 
J.H. Bradbury, W.D. William^ 
corner is marked additionally by another two 
'hadziid' setae, the palm and dactyl are more 
spinous, the dactyl lacking recumbent inner teeth. 
The oostegites bear more distal setae, urosomite 
one has three rather than two ventro-distal spines 
at the base of the first uropod, the peduncles of the 
first uropod are less spinous, the second article of 
the third uropod and telsonic lobes are more 
slender, the telson lacks penicillate setules. 
Liagoceradocus subthalassicus is similar to L. 
dentiferus Ledoyer, 1982 in that the third article of 
the maxilliped is curved and the length less than 
twice the width, second and third coxal plates are 
longer than wide, and the palm of the second 
gnathopod bears both setae and spines. L. 
subthalassicus differs from L. dentiferus in the 
absence of proximal spines on the medial margins 
of the telsonic lobes, such spines being located in 
the distal half of the margin, the width of the fourth 
coxa is as great as the length, and the second article 
of pereopods 5 to 7 is post-ventrally lobate. 
Liagoceradocus subthalassicus differs from L. acutus 
Andres, 1978 in the greater relative length of the 
third article of the peduncle of the first antenna, 
shorter flagellum with fewer articles and 
aesthetascs on both antennae, rudimentary inner 
lobes to the lower lip, on the mandible an 
extremely short molar seta, simple right lacinia 
mobilis, fewer setate accessory spines, short first 
article of the palp which bears an extra E seta on 
the third article, fewer denticulate spines on the 
outer lobe of the first maxilla which bears 
asymmetrical palps, fewer more robust setae in the 
oblique facial row of the second maxilla, the inner 
lobe of the maxilliped without a sub-distal oblique 
row of setae rather with a sparse sub-marginal 
facial row. The coxa and second article of the first 
gnathopod have fewer spines and setules, the 
palmar angle distinct on the right side only 
although marked by fewer spines, the palm is 
without spinules and with few setae. The coxa and 
second article of the second gnathopod are less 
armed, the lobe of article five is keel like, the palm 
bears two lateral and two medial spines, the 
palmar margin (of the female) is equal to the free 
posterior margin, coxae of pereopods 3 and 4 are 
equal, coxa four bearing a small posterior seta, 
article two of P5 is more expanded and ovate, and 
there are few armaments on any article. Pereopod 
6, article one is equal to that of P5. Coxal gills 3-4 
are largest, gill 6 smallest. The epimera are without 
an oblique ridge and bear few spines; E2-3 with 1- 
2 antero-sub-marginal spines, and the posterior 
comers of each is rounded rather than acuminate. 
The pleopods are without bifid spines. The first 
urosomite bears three ventro-distal spines at the 
base of the first uropod, the peduncle of the second 
uropod bears two apico-medial spines and a disto- 
medial group of four small spines, the outer ramus 
has three dorso-facial rather than two medio-dista] 
spines, the apex of the third uropod an additional 
sub-apical setule. The telson is lanceolate rather 
than ovate, bearing one rather than two disto- 
lateral spines and is without penicillate setules. 
L. subthalassicus differs from L. branchialis in: thq 
absence of a distinct posterior lobe on the carpus of 
the second gnathopod; reduced rather thaq 
moderate molar seta; a greater number of medial 
setae on the second and third articles of thq 
mandibular palp; longitudinal rather than obliqutj 
row of medial plumose setae on the inner plate of 
the first maxilla; greater pubescence of thq 
maxillae; denticulate rather than spinous apical 
setae on the L outer plate of maxilla 1; absence of q 
small post-ventral lobe on the second article of P5; 
three ventro-distal spines rather than a small, 
single seta on urosomite 1 at the base of Ul; 
presence of accessory retinacula on pleopods 1-3; 
greater setation of Ul-3; presence of lateral 
setation, but absence of penicillate setules on th<> 
telson which is not tumid laterally. 
Etymology 
Named for the nature of the type locality. 
Liagoceradocus branchialis sp. nov 
Figures 4—6 
Material Examined 
Holotype 
3 'a' from sample BES 4282, body length 4.0 mm. 
Cave C-28, Cape Range peninsula. Western 
Australia, Australia (22°25'S, 113°46'E), 26 May 
1995, in saline water, A.A. Poole, D. Warren (WAM 
257-95). 
Para types 
1 3 ‘b’ from sample BES 4282, body length 
3.5mm, same data as holotype (WAM 272-95); 3 3 
same data as holotype except 24 May 1995, body 
length 3-5mm (WAM 273-95 to 275-95); 1 3, same 
data as holotype except 25 May 1995, body length 
3.5 mm (WAM 256-95). 
Type locality 
Bundera Cenote, (C-28) below pycnocline in a 
tidal, almost anoxic, anchialine cave (22°25'S, 
113°46'E) on North West Cape, Western Australia. 
Diagnosis 
Pereonites and pleonites bearing few fine dorsal 
and dorso-lateral setae, without carinations or 
teeth. Head. Rostrum weak, lateral cephalic lobes 
weakly projecting, antennal sinus small and 
distinct, eyes absent. First antenna elongate, longer 
than A2, ratio of peduncular articles 24:27:10, 
