3- J 
_ —U- - _ D 
_ —- — ~~ U 
i- 1 - 1 - 1 -r 
-3-2-10 1 2 
Function 2 
Figure 4 Plots of Functions 1 and 2 from Canonical 
Variate Analysis among each island 
population and the Peninsula Malaysia 
populations of Hipposideros bicolor and based 
on a subset of five external characters (see 
text). Population codes as for Figure 2. 
Group from the Selaru Group and partially 
separated it from the Timor Group; it also partially 
separated the Timor and Roti/Savu Groups from 
the Selaru Group (Figure 5). The characters with 
high coefficients (>0.5) on Function 2 were digit 3 
phalanx 1, ear length and forearm length. Function 
3 (12.6% of variance) also separated the Malay 
Peninsula Group from the Roti/Savu Group. The 
characters with high coefficients (>0.5) on Function 
3 were digit 4 and 5 metacarpal lengths (Table 3b). 
In summary, the above statistical analyses 
indicated that the specimens of H. bicolor from the 
Lesser Sunda islands differed morphologically in 
both cranial, dentary, dental and external body 
characters from H. bicolor atrox from Peninsula 
Malaysia and also on cranial, dentary and dental 
characters from H. b. bicolor from Java. However, 
among this island chain the populations were 
further differentiated on cranial, dentary and 
dental characters into three clear forms (Timor, 
Selaru and Roti/Savu). The two Sumbawa island 
specimens grouped closely with those from Selaru 
island on external body morphology but on cranial, 
dentary and dental morphology the single 
specimen available for study was not closely 
associated with any of the other specimens. The 
Sumba island specimen is clearly closely associated 
with the Selaru form on cranial, dentary and dental 
characters and is tentatively referred to that form. 
The Sumbawa specimens, described in detail by 
Kitchener and Maharadatunkamsi (1995), are not 
D.J. Kitchener, Y. Konishi, A. Suyanto 
Function 2 
Figure 5 Plots of Functions 1 and 2 from Canonical 
Variate Analysis among four population 
groupings (Selaru, Timor, Roti/Savu and 
Peninsula Malaysia) - with the few Sumba, 
Sumbawa and Java specimens unallocated - 
and based on a subset of five external body 
characters. Population codes as for Figure 2. 
referred to any of these Lesser Sunda subspecies. 
These two specimens agree in size with the 
measurements of a single specimen from Ruteng, 
Flores island reported in Hill and Rozendaal 
(1989). Further specimens will probably allow 
recognition of the Sumbawa and Flores form as a 
unique subspecies. 
The following section formally recognises three 
new subspecies of H. bicolor in the Lesser Sunda 
islands. 
SYSTEMATICS 
Hipposideros bicolor tanimbarensis subsp. nov., 
Kitchener 
Holotype 
Western Australian Museum (WAM) No. 
M44287; adult female; carcass fixed in 10% 
formalin and preserved in 70% ethanol; skull 
separate; collected by Ir. Ibnu Maryanto (Museum 
Zoologicum Bogoriense) and Ron Johnstone, 
Western Australian Museum), on 25 April 1993. 
Type locality 
Wesuri Cave, near Adaut, Selaru island, 
Tanimbar Island Group (8°09'S, 131°08'E); at sea 
level. 
Paratypes (all from type locality) 
122 2, 12dd; WAM (M43818, M44248-9, 
