Morphological variation in Hipposideros bicolor 
M44252, M44254, M44259, M44261-8, M44271-2, 
M44275-6, M44279, M44281-3, M44286, M44290). 
Diagnosis 
Hipposideros bicolor tanimbarensis differs from H. 
b. bicolor [including also measurements from Hill 
(1983) and Tate (1941)] in averaging smaller in 
most cranial, dentary and dental measurements, 
except for tympanic bulla length and intercochlear 
distance. For example: absolutely smaller greatest 
skull length 17.2-18.0 (N = 18) v. 18.2-19.0 (3) and 
least interorbital breadth 2.5-2.7 (18) v. 2.9-3.0 (4). 
Posterior noseleaf breadth larger 5.1-5.9 (24) v. 4.7- 
4.9 (2). Interlachrymal distance narrower relative 
to dentary length (Figure 6). Dorsal pelage darker, 
tipped with Fuscous rather than Burnt Umber. 
H. b. tanimbarensis differs from H. b. atrox in 
having a narrower: zygomatic width 8.8-9.3 (18) v. 
9.5-9.7 (3); least interorbital breadth 2.5-2.7 (18) v. 
2.8-2.9 (3); M 3 M 3 width 5.5-6.0 (17) v. 6.1-6.4 (3). It 
has a longer digit 5 metacarpal length relative to 
forearm length (Figure 7); and narrower least 
interorbital breadth relative to dentary length 
(Figure 8). Gians penis distal end sharper and less 
rounded in craniocaudal view, and from lateral 
view distal tip longer (Figure 9). Baculum more 
gracile, base not as broad and not bilobed; arms of 
distal bifurcation not as widely spaced or as broad 
(Figure 10). Dorsal pelage darker, tipped with 
Fuscous rather than Cinnamon. 
H. b. tanimbarensis differs from H. b. hilli subsp. 
nov. in averaging larger in all characters, except 
least interorbital breadth, interlachrymal distance, 
cochlea length, and digit 5 metacarpal length. 
Figure 6 Plot of interlachrymal distance versus dentary 
length for populations of Hipposideros bicolor. 
Population codes as for Figure 2. 
187 
Forearm length (mm) 
Figure 7 Plot of digit 5 metacarpal length versus 
forearm length for populations of 
Hipposideros bicolor. Population codes as for 
Figure 2. 
Dentary length shorter relative to least interorbital 
breadth (Figure 8). Dorsal pelage darker, tipped 
with Fuscous rather than Burnt Umber. 
H. b. tanimbarensis differs from H. b. selatan subsp. 
nov. in having a narrower interlachrymal distance 
4.4-4.6 (18) v. 4.7-4.9 (7) and narrower least 
interorbital breadth relative to dentary length 
(Figure 8). Dorsal pelage darker, tipped with 
Fuscous rather than Burnt Umber. 
Description 
A moderate sized H. bicolor subspecies with 
greatest skull length 17.66 (17.16-17.98) 18 and 
forearm length 42.9 (40.3-45.0) 25. For other 
measurements see Table 2a, b. It agrees closely 
with the general description of the cranium, 
dentition and external body characters of this 
species as described by Hill (1963). Hipposideros b. 
tanimbarensis has zygomata broad, occasionally 
massive, with superior jugal projection low; 
zygomatic width narrower than mastoid width 
9.08 (8.75-9.26) 18 v. 9.21 (9.03-9.41) 18; least 
interorbital breadth narrow 2.58 (2.45-2.71) 18; 
crown area of outer lower incisor slightly larger 
than that of inner lower incisors; anterior lower 
premolar three-quarters length and half to three- 
quarters height of posterior lower premolar; and 
vomer projection into mesopterygoid fossa only 
slightly thickened. 
Pelage of ventral surface Smoke Gray; dorsal 
surface Smoke Gray base tipped with Fuscous. The 
patagia Dusky Brown. 
