188 
D.J. Kitchener, Y. Konishi, A. Suyanto 
Least interorbital breadth 
Figure 8 Plot of dentary length versus least interorbital 
breadth for populations of Hipposideros 
bicolor. Population codes as for Figure 2. 
Penis ca. 7 mm long. Gians penis with distal head 
of variable shape; in craniocaudal view it ranges 
from a blunt rod shape (Figure 9b) to an arrow 
shape (Figure 9c) to a broadly rounded head with 
median distal slit (Figure 9d). Baculum ca. 3.0-3.2 
a b 
mm long; base slightly broadened, long narrow 
shaft with short bifurcated distal tip (Figure 10b,c). 
Distribution 
Known only from the type of locality at Wesuri 
Cave, near Adaut, Selaru island, Tanimbar Group. 
Etymology 
Named after the Tanimbar Group of islands 
which it inhabits. 
Referred Specimen 
A single adult female (WAM M30345) mist 
netted over a river in a large limestone cave at 
Waikelosawah, Sumba island (9°36 , S,119°29'E) is 
tentatively referred to H. b. tanimbarensis; its 
measurements are presented in Tables 2a, b. On 
the basis of the more important cranial, dental and 
dentary measurements the DFA allocated it to H. b. 
tanimbarensis. Its glans penis is similar to Figure 9c; 
its baculum has a length of 2.74 mm and is similar 
to Figure 10b. 
Hipposideros bicolor hilli subsp. nov. Kitchener 
Holotype 
Western Australian Museum (WAM) No. 
M34953; adult male; carcass fixed in 10% formalin 
and preserved in 70% ethanol; skull separate; 
c d 
Figure 9 Gians penis of (a) Hipposideros bicolor atrox from Peninsula Malaysia (WAM M21159); (b) and (c) H. b. 
selatan subsp. nov. from Roti island (WAM M35375 and WAM M35507, respectively) and (d) H. b. hilli 
subsp. nov. from Timor island (WAM M34959). From (i) cranial, (ii) lateral and (iii) caudal views. Scale 
lines, 0.5 mm. 
