Morphological variation in Hipposideros bicolor 
189 
collected on 17 October 1990 by Dr Ken Aplin 
(Western Australian Museum) and Bapak Boeadi 
(Museum Zoologicum Bogoriense). 
Type Locality 
Mist netted in a cave at Oimoro, near Panite, 
West Timor, Nusa Tenggara Timur, Indonesia 
(9°50'S, 124°29'E); at sea level. 
Paratypes 
From the following localities on West Timor: 
Baumata (10°11'S, 123°43'E), 72 9, 56 6, WAM 
M(30060, 30098-9, 30107, 30110, 30115-6, 30118-22; 
Bauraen (10°15'S, 123°43'E), 12, WAM M35010; 
Panite (9°50'S, 124"29'E), 112 2 9 6 6, WAM 
M(34889-92, 34894, 34951-2, 34954-6, 34958-9, 
34961, 34963-5, 34967-8, 34973-4). 
Diagnosis 
Hipposideros b. hilli differs from H. b. bicolor 
[including also Hill (1983) and Tate (1941) 
measurements] in averaging smaller in most 
measurements, except tympanic bulla length, M 3 
breadth, ear length and digit 5 metacarpal length. 
For example, absolutely smaller in both greatest 
skull length 16.8-17.8 (27) v. 18.2-19.0 (3) and 
forearm length 40.8-44.4 (34) v. 44.6-46.8 (6). 
Dentary length relative to least interorbital breadth 
shorter (Figure 8). 
Hipposideros b. hilli has similar cranial and 
external body measurements to H. b. atrox but 
differs in having considerably smaller dental 
measurements (see Table 2a). Dentary length 
relative to both interlachrymal distance and least 
interorbital breadth shorter (Figures 6 and 8, 
respectively). Digit 5 metacarpal length longer 
relative to forearm length (Figure 7). Gians penis 
distal end less rounded in craniocaudal view and 
from lateral view distal tip longer (Figure 9). 
Baculum more gracile, base not as broadened and 
not bilobed, arms of the distal bifurcation not as 
widely spaced or as broad (Figure 10). Dorsal 
pelage darker, tipped with Burnt Umber rather 
than Cinnamon. 
Hipposideros b. hilli differs from H. b. tanimbarensis 
as described in the earlier diagnosis of that 
subspecies. 
Hipposideros b. hilli differs from H. b. selatan in 
averaging smaller in most cranial, dental and 
dentary characters. For example, greatest skull 
length 17.3 (16.8-17.8) 27 v. 17.6 (17.4-17.9) 7. 
Interlachrymal distance and dentary length 
generally smaller (Figure 6) and dentary length 
relative to least interorbital breadth generally 
smaller (Figure 8). 
Description 
Hipposideros b. hilli is on average, the smallest 
i i 
Figure 10 Baculum of (a) Hipposideros bicolor atrox 
from Peninsula Malaysia (WAM M21158), 
(b) H. b. selatan subsp. nov. from Roti island 
(WAM M35507), and (c), H. b. hilli subsp. 
nov. from Timor island WAM M34959). 
From (i) caudal, (ii) lateral and (iii) cranial 
views. Scale lines 0.5 mm. 
form of H. bicolor in the Lesser Sunda Islands. It 
has a greatest skull length of 17.29 (16.82-17.78) 27 
and forearm length of 42.3 (40.8-44.4) 34. Other 
measurements of cranium, dentition, dentary and 
external body characters (see Table 2a, b) are 
similar to H. b. tanimbarensis. Its zygomatic width 
is much narrower than the mastoid width 8.77 
(8.49-8.96) 27 v. 9.10 (8.73-9.31) 27. 
