New cuttlefishes from the North West Shelf 
281 
Table 2 Sepia plana sp. nov.; measurements (mm), counts and indices of 1 male (M) and 6 females (F), 
Museum 
MV 
MV 
MV 
MV 
MV 
MV 
MV 
Reg. No. 
F77206 
F80339 
F67700 
F77205 
F67700 
F58848 
F77204 
(Holotype) 
(Paratype) 
(Paratype) 
(Paratype) 
(Paratype) 
(Paratype) 
(Paratype) 
Sex 
M 
F 
F 
F 
F 
F 
F 
Maturity 
mature 
immature 
mature 
mature 
mature 
mature 
mature 
ML 
99.3 
63.0 
84.4 
100.0 
127.1 
148.7 
151.2 
MWI 
57.4 
61.6 
63.3 
57.4 
55.1 
57.9 
54.8 
AMHI 
16.8 
14.4 
17.2 
17.3 
14.3 
12.8 
12.2 
VMLI 
90.8 
82.2 
78.4 
80.6 
87.6 
94.4 
93.6 
FWI 
13.8 
12.2 
- 
10.9 
7.4 
8.9 
9.3 
Flla 
14.4 
15.6 
- 
8.9 
16.4 
10.7 
14.9 
Flip 
9.3 
10.0 
- 
9.9 
13.0 
10.5 
14.4 
FuLI 
38.3 
35.1 
35.7 
32.0 
28.7 
28.6 
34.4 
FFul 
16.1 
14.3 
14.8 
13.0 
11.4 
17.8 
15.5 
HLI 
26.3 
40.5 
- 
23.2 
- 
27.6 
- 
HWI 
48.8 
50.8 
- 
46.3 
48.4 
44.4 
39.6 
EDI 
12.1 
17.3 
- 
16.1 
12.4 
12.4 
- 
ALII 
48.8 
44.4 
- 
35.5 
33.8 
38.0 
28.8 
ALI2 
40.8 
41.3 
- 
44.5 
- 
44.0 
36.4 
ALI3 
41.3 
41.3 
- 
37.5 
39.7 
40.3 
36.4 
ALI4 
45.8 
41.3 
- 
39.5 
41.3 
40.0 
38.4 
ASInl 
1.61 
1.27 
- 
1.07 
1.08 
1.26 
0.89 
ASIn2 
1.88 
1.27 
- 
1.12 
0.98 
1.31 
0.99 
ASIn3 
1.51 
1.35 
- 
1.00 
0.90 
1.21 
1.04 
ASIn4 
1.33 
1.27 
- 
1.00 
0.79 
1.26 
0.96 
ASCI 
94 
102 
- 
128 
102 
- 
116 
ASC2 
90 
110 
- 
136 
- 
- 
146 
ASC3 
112 
130 
- 
145 
150 
- 
142 
ASC4 
118 
146 
- 
160 
172 
- 
145 
ASInl4 
1.26 
- 
- 
- 
- 
_ 
_ 
ASInl4m 
0.76 
- 
- 
- 
- 
- 
- 
C1L1 
19.1 
15.7 
- 
15.5 
18.5 
15.5 
_ 
CIRC 
8 
10 
- 
8 
- 
8 
_ 
TrRC 
36 
34 
- 
36 
45 
36 
- 
C1SI 
0.22 
0.40 
- 
0.27 
0.29 
0.34 
_ 
CISId 
0.22 
0.40 
- 
0.25 
0.31 
0.27 
_ 
CISIv 
0.22 
0.32 
- 
0.27 
0.29 
0.27 
_ 
GiLC 
29 
25 
- 
30 
28 
28 
_ 
GiLI 
- 
34.9 
31.1 
37.9 
42.0 
37.0 
_ 
SpLI 
8.6 
- 
- 
- 
- 
- 
- 
SpWI 
4.7 
- 
- 
- 
- 
- 
EgDI 
- 
- 
4.0 
- 
8.5 
10.0 
CbL 
- 
- 
84.3 
98.2 
126.9 
- 
_ 
CbWI 
- 
- 
45.4 
45.8 
42.9 
_ 
- 
SLI 
- 
- 
0.6 
1.4 
0.4 
_ 
_ 
StZI 
- 
- 
50.9 
57.0 
50.7 
_ 
_ 
LoLI 
- 
- 
45.8 
42.9 
48.5 
_ 
_ 
LoL/StZ (%) 
- 
- 
90.0 
75.2 
95.6 
- 
- 
approximately half funnel length; FFul male 16.1, 
females 11.4- 14.5 -17.8 (SD, 2.5). Funnel organ 
dorsal elements thick, inverted V-shape; ventral 
elements oval (Figure 3A). Mantle-locking cartilage 
curved, with semicircular ridge; funnel-locking 
cartilage with depression which corresponds to 
ridge (Figure 3B). Head short; HLI male 26.3, 
females 23.2- 25.4 27.6 (SD, 3.1); broad, narrower 
than mantle; HWI male 48.8, females 39.6 —14.7 - 
48.4 (SD, 3.7). Eyes large; EDI male 12.1, females 
12.4 -13.7- 16.1 (SD, 2.1); ventral eyelids present 
(Figure 2A). 
Male and female arms subequal in length (Table 
3). Arm length index of longest arms in male 48.8, 
females 36.4- 41.6 -44.5 (SD, 4.6). Protective 
membranes (both sexes) narrow; normal, not 
thickened. Distal arm tips (both sexes) strongly 
attenuate, suckers enclosed by protective 
membranes. Arm suckers tetraserial in both sexes. 
Suckers in males normal in size (not greatly 
enlarged); larger than female arm suckers (Table 
3). Inner ring of chitinous sucker rims with blunt 
projections; infundibulum with 12-14 rows of 
ovoid-hexagonal processes without pegs. 
