New cuttlefishes from the North West Shelf 
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Figure 4 A-D, Sepia plana sp. nov.; A, arm 2 sucker rim, female paratype MV F77205, 100 mm ML, scale bar 100 pm; 
B, club sucker rim, female paratype MV F67700, 127.1 mm ML, scale bar 40 pm; C, radula, female paratype 
MV F77205, 100.0 mm ML, scale bar 500 pm; D, enlargement of 1st laterals and rhachidian (centre) teeth, 
same specimen, scale bar 200 pm. 
elongate, rectangular; at periphery processes 
smaller, sub-rectangular (Figure 4B). Swimming 
keel of club extends well beyond carpus (Figure 
5A). Dorsal and ventral protective membranes not 
fused at base of club; joined to stalk; dorsal and 
ventral membranes same length, extend beyond 
carpus along stalk; approximately equal width; 
dorsal membrane forms shallow cleft at junction 
with stalk. 
Gills with 28-30 lamellae per demibranch; GiLC 
male 29, females 28-29-30 (SD, 1). Gill length: GiLI 
females 31.1-37.0-42.0 (SD, 4.5). 
Buccal membrane without suckers. Upper beak 
(Figure 5B) rostrum bluntly pointed, length greater 
than width; cutting edge slightly curved; hood high 
above crest posteriorly; lateral wall shallowly 
indented posteriorly; wings and hood narrow, and 
short; jaw angle approximately 90°; hood dark 
brown, lighter on posterior edge, crest darkly 
pigmented. Lower beak (Figure 5C,D) rostrum 
bluntly pointed, protrudes only slightly; cutting 
edge curved; hood low on crest; crest straight, no 
indentation on lateral wall edge; hood and wings, 
width narrow; hood notch deep, broad (Figure 5D); 
wings widely spaced; crest wide with deep notch; 
rostrum pigmented dark brown, fading gradually 
on wing, crest pigmented, darkest medially fades 
on lateral walls. Radula with 7 teeth per row; 
homodont; rhachidian teeth narrow, not broad 
based, slender, triangular, sides straight (Figure 
4C,D); first lateral teeth same length as rhachidian 
teeth, asymmetrical with mesocone displaced 
toward centre of radula, heel narrow (Figure 
4C,D); second laterals longer than first, mesocone 
displaced towards centre, heel broad; marginal 
teeth elongate, tapered, curved, rounded 
proximally and strongly indented (Figure 4C). 
Digestive tract (Figure 5E): paired salivary glands 
approximately one-third length of buccal mass; 
paired digestive glands large, located close 
together, with sub-triangular lobes posteriorly, 
ducts connect digestive glands near midline with 
caecum (not shown in figure), ducts with branched 
attached pancreatic tissue; oesophagus runs 
dorsally along median junction of digestive glands, 
joins sac-like stomach immediately posterior to 
digestive glands; caecum disc-like, grooved in 
blunt V-shape anteriorly, surface lining finely 
pleated; intestine undifferentiated; ink sac and anal 
flaps well developed. 
Male reproductive tract: testis on left posterior 
side of viscero-pericardial coelom; at distal end, 
convoluted vas deferens opens into broad cone- 
shaped mucilaginous gland, then narrower, curved 
spermatophoric gland (Figure 6A). Close to 
junction with lobe-shaped accessory gland and 
