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Figure 5 A-E, Sepia plana sp. nov.; A, tentacular club, female paratype MV F7720S, 100.0 mm ML, scale bar 2 mm; B, 
upper beak, lateral view; C, lower beak, lateral view; D, lower beak, ventral view (B-D, male holotype MV 
F77206, 99.3 mm ML, scale bar 3 mm); E, digestive tract, female paratype (MV F80339), 63.0 mm ML, scale 
bar 6 mm, (A - anus; BM - buccal mass; C - caecum; DG - digestive gland; IS - ink sac; P - pancreas; S - 
stomach; SG - salivary gland). 
gland appendix, delicate ciliated canal joins 
spermatophoric gland; distal deferent canal 
connects appendix of accessory gland to 
spermatophore storage sac; genital orifice opens 
dorsal to left gill in anterior end of mantle cavity. 
Spermatophores: cement body bipartite (Figures 
6B, 7); aboral end elongate, cylindrical, connects to 
sperm reservoir via narrow duct which extends 
from tip of cement body; oral end of cement body 
approximately half length of aboral end, similar 
width; connects to aboral end via narrow neck; 
middle tunic commences along aboral part of 
cement body; ejaculatory apparatus coiled, extends 
into oral dilation of spermatophore. 
Spermatophores 8.5 mm long, 0.4 mm wide; SpLI 
8.6; SpWI 4.7. 
Female reproductive tract: ovary hangs from 
dorsal wall of posterior viscero-pericardial coelom. 
Oviduct thin-walled, continuous with body cavity; 
distally with thickened, glandular walls (oviducal 
glands). Nidamental glands in mature animals 
occupy large portion of ventral side of mantle 
cavity. Accessory nidamental glands anterior to 
nidamental glands (Figure 6C). Eggs spherical, 4.0- 
7.5-10.0 mm diameter (SD, 3.1); EgDI 4.0-5.4-6.6 
(SD, 1.3). 
