New cuttlefishes from the North West Shelf 
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Figure 6 A-D, Sepia plana sp. nov.; A, male reproductive tract (testis not shown), holotype MV F77206, 99.3 mm ML, 
scale bar 1 mm, (AAG - appendix of accessory gland; AG - accessory gland; CC - ciliated canal; DDC - 
distal deferent canal; GO - genital orifice; MG - mucilaginous gland; SG - spermatophoric gland; SSS - 
spermatophore storage sac; VD - vas deferens); B, spermatophore, holotype MV F77206, 99.3 mm ML, scale 
bar 1 mm; C, female reproductive tract, paratype MV F77205, 100 mm ML, scale bar 20 mm, (ANG - 
accessory nidamenta! gland; GO - genital orifice; NG - nidamental gland; O - ovary); D, posterior end of 
cuttlebone, lateral view, female paratype MV F67700, 84.4 mm ML, scale bar 1 mm. 
Subdermal cartilaginous layer between 
cuttlebone and skin absent. Cuttlebone length 
approximately equal to mantle length; outline oval 
(Figure 8A,B); CbL females 84.3- 103.1 -126.9 (SD, 
21.7); CbWI females 42.9-44.7-45.8 (SD, 1.6). 
Cuttlebone bluntly rounded anteriorly; acuminate, 
acute posteriorly (Figure 8A,B); recurved ventrally 
(Figure 6D). Dorsal surface creamy white; flat 
medially, flat laterally (Figure 8A); granulose 
(Figure 8C). Dorsal median rib present, indistinct, 
broadens anteriorly; lateral ribs present, distinct. 
Chitin borders lateral margins of cuttlebone. Spine 
present, short (Figures 6D, 8A,C); SLI females 0.4- 
0.8-1.4 (SD, 0.5); curves dorsally (Figure 6D); 
keel(s) absent; cuttlebone smooth between spine 
and outer cone; ventral notch at base of spine 
absent. Dorso-posterior end of cuttlebone smooth, 
without median longitudinal ridge anterior to 
spine. Striated zone flat; StZI females 50.7 -52.9- 57 
(SD, 3.6). Last loculus flat; LoLI females 42.9 -45.7 - 
48.5 (SD, 2.8); slightly shorter than striated zone, 
LoL/StZ(%) females 75.2- 86.9 -95.6 (SD, 10.6). 
Sulcus absent. Anterior striae broad inverted U- 
shape (Figure 8B). Limbs of inner cone extend 
anteriorly to end of striated zone. Inner cone limbs 
narrow, broaden very slightly posteriorly, not 
raised to form ledge posteriorly; not thickened 
posteriorly. Outer cone calcified; narrow 
