302 
C.C. Lu, A. Reid 
Figure 20 A-C, Sepia sulcata Hoyle; A, hectocotylised left ventral arm, holotype BMNH 89.4.24.77, 53.2 mm ML; B, 
modified portion of hectocotylized left ventral arm, MV F56919, 67.6 mm ML; C, right ventral arm, male 
MV F56919, 67.6 mm ML. Scale bars 2 mm. 
wings and hood narrow and short; lateral wall 
shallowly indented posteriorly; jaw angle 
approximately 90°; hood darkly pigmented, fades 
toward posterior margin; crest pigmented along 
midline, lateral walls pale. Lower beak cutting 
edge straight; hood low on crest (Figure 21C); crest 
straight; lateral wall edge without indentation; 
hood and wings, width narrow; hood notch deep 
(Figure 21D); crest wide; jaw angle obtuse; hood 
darkly pigmented, fades toward posterior margin; 
crest pigmented along midline, lateral walls pale. 
Radula with 7 teeth per row; homodont; rhachidian 
teeth with broad, strongly indented bases, narrow, 
tapered, blunt distally (Figure 19D,E); first lateral 
teeth slightly longer, broader than rhachidian teeth, 
with wide heels and bases strongly indented, 
asymmetrical with mesocone displaced toward 
centre of radula (Figure 19D,E); second laterals 
similar in shape, slightly longer than first laterals; 
marginal teeth elongate, curved, bluntly rounded 
distally, with rounded heel, strongly indented 
proximally (Figure 19D). Digestive tract (Figure 
21E): paired salivary glands approximately one- 
third length of buccal mass; paired digestive glands 
large, located close together, with sub-triangular 
lobes posteriorly, ducts (not shown in figure) 
connect digestive glands near midline with 
caecum, with branched attached pancreatic tissue; 
oesophagus runs dorsally along median junction of 
digestive glands, joins sac-like stomach 
immediately posterior to digestive gland; caecum 
disc-like, grooved in blunt V-shape anteriorly, 
surface lining finely pleated; intestine 
undifferentiated; ink sac and anal flaps well 
developed. 
Male repoductive tract: testis on left posterior 
side of viscero-pericardial coelom; at distal end, 
convoluted vas deferens (Figure 22A) opens into 
broad, cone-shaped mucilaginous gland, then 
narrower, curved, spermatophoric gland. Close to 
junction with lobe-shaped accessory gland and 
gland appendix, delicate ciliated canal joins 
spermatophoric gland; distal deferent canal 
connects appendix of accessory gland to 
spermatophore storage sac; genital orifice opens 
dorsal to left gill in anterior end of mantle cavity. 
