306 
C.C. Lu, A. Reid 
Figure 23 Sepia sulcata Hoyle; spermatophore, oral end, 
MV F56919, 67.6 mm ML, scale bar 1 mm. 
acute anteriorly; acuminate, acute posteriorly 
(pointed). Dorsal surface yellowish; flat medially, 
curved, convex laterally; texture smooth, not 
pustulose. Dorsal median rib present; distinct; 
broadens anteriorly; lateral ribs present, distinct. 
Chitin present over entire surface of cuttlebone, 
with thicker band on each side. Spine present; 
short; SLI males 2.5—3.3-4.4 (SD, 0.8), females 1.2— 
2.9-3.5 (SD, 0.8); curves dorsally; keel(s) absent; 
cuttlebone smooth between spine and outer cone; 
ventral notch at base of spine absent. Dorso- 
posterior end of cuttlebone with short, median 
longitudinal ridge anterior to, and separate from 
spine (Figure 24A). Striated zone convex (slightly); 
StZI males 60-63.5-67.8 (SD, 2.8), females 57-64— 
71.3 (SD, 5.1). Last loculus flat; LoLI males 28.9- 
35.0-40.3 (SD, 3.9), females 25-33.7-39.2 (SD, 6.1); 
one-third length of striated zone (approximately); 
LoL/StZ(%) males 44.0-55.4-67.2 (SD, 8.4), females 
35- 53.5- 66.5 (SD, 13.1). Sulcus extends along entire 
length of bone; shallow, narrow; flanked by 
rounded ribs bordered laterally by shallow grooves 
(distinctive). Anterior striae inverted U-shape 
(slightly wavy due to ribs). Limbs of inner cone 
extend anteriorly to end of striated zone. Inner 
cone limbs broaden posteriorly, raised to form 
ledge; ledge flat, not thickened (Figures 24B, 25). 
Outer cone chitinous, not calcified; narrow 
throughout; lateral limbs not flared ventro- 
laterally; outer cone strongly recurved ventral to 
spine, forms cup-like structure. 
Body papillae present; dorsal mantle with 
longitudinal row of up to 10 short longitudinal 
ridges along each side close to fins (not visible in 
all specimens examined); ventral mantle with 
longitudinal row of 6 narrow ridges along each 
side close to fins; anterior-most pair and posterior 
2 pairs shorter than rest (Figure 18C) (short ridges 
sometimes difficult to detect in small specimens). 
Head and arm papillae absent. Colour (alcohol 
preserved specimens): Head pale, without colour 
pattern. Dorsal mantle pale buff pinkish-brown; 
paired dorsal eye spots absent. Fins pale; without 
markings at base. Arms without markings. Ventral 
pigment absent. Dorsal and ventral longitudinal 
ridges pinkish-orange. 
Original reference 
Hoyle, W.E. (1885). Diagnoses of new species of 
cephalopoda collected during the cruise of H.M.S. 
"Challenger." Part II. The Decapoda. Annales and 
Magazine of Natural History (5) 110:181-203. 
Type locality 
Indonesia: Arafura Sea, off Ki Is., 5°49'15"S 
132°14'15"E; 255 m. 
TYPE: Holotype, BMNH 89.4.24.77, male, 53.2 
mm ML. 
Distribution 
Indonesia, Arafura Sea, off Ki Is., 5°49'15"S 
132°14'15"E to Australia: Western Australia, North 
West Shelf, 19°58'S 115°13'E; depth range 404-150 
m. Habitat mud (holotype blue mud), silt, rock. 
Remarks 
This species previously was known only from the 
