Eocene cranioid brachiopod 
315 
Remarks 
In overall size Westralicrania zenobiae is generally 
larger than Westralicrania allani (4.8-11.7 mm for 
IV. zenobiae compared with 2.2-6.8 mm for W. 
allani). The posterior muscle scars are 
comparatively larger (20% : 12.5% of total valve 
width), whereas the median septum is 
comparatively shorter in IV. zenobiae than W. allani 
(11.1% : 28.6% of total valve length). There are 
significantly more growth lines in the 
pseudointerarea of IV. zenobiae than in IV. allani as 
it is comparatively longer. The spines on the 
exterior surface of IV. zenobiae are relatively shorter 
on IV. zenobiae, but they radiate in lines whilst those 
on W. allani have no apparent regularity. The 
pseudointerarea is anacline in W. zenobiae and 
apsacline in W. allani. 
W. allani was originally described from Shark 
Bay (Cockbain 1966). Nine further specimens of W. 
allani (WAM 84.579[x4], 84.580 - 84.583, 92.714) 
have been collected from the Cardabia Calcarenite, 
Giralia Range, Carnarvon Basin, Western Australia 
of Middle Paleocene age. They accord well with 
the description of the species by Cockbain (1966). 
The size range of the specimens is slightly larger 
(6.31-8.76 mm) than those from Shark Bay. 
ACKNOWLEDGEMENTS 
I would like to thank Dr J. Backhouse for the use 
of the type specimens of Westralicrania allani from 
the Geological Survey collection, Ms K. Brimmell 
for photographing the specimens of Westralicrania 
zenobiae and my wife Zenobia for the camera lucida 
diagram of the ventral shell. Special thanks must 
go to Dr K. J. McNamara for all his assistance, 
advice and support in preparing this paper. 
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Manuscript received 14 February 1997; accepted 4 June 1997. 
