334 
SYSTEMATICS 
Family Geophilidae 
Genus Tuoba Chamberlin 
Tuoba Chamberlin, 1920: 35. 
Nesogeophilus Verhoeff, 1924: 413. Synonymised by 
Attems, 1929:184. 
Algerophilus Brolemann, 1925: 250. Synonymised by 
Brolemann, 1925: 250. 
Type species 
Tuoba: Tuoba curticeps Chamberlin, 1920 (from 
Solomon Islands). 
Geophilus ( Nesogeophilusy. Geophilus 
(Nesogeophilus) b'ackstromi Verhoeff, 1924 (from Juan 
Fernandez). 
Algerophilus: Geophilus hispanica Meinert, 1870 
(from Spain, Seville, Granada). 
Diagnosis 
Clypeus bounded by broad paraclypeal sutures, 
labrum tripartite. Prosternum with chitin lines. 
Carpophagus structures occupy the segments to 
the transition (Eason, 1964). Coxopleural glands are 
complex, with each coxopleuron having a single 
large, concealed glandular crypt that is 
multiglanduiar and multicanaliculate. The 
pretarsal anterior parunguis is spiniform, greatly 
elongate and much longer than the minute 
posterior parunguis (Figures 3, 15, 29, 40). Anal 
claw present. Littoral. 
Tuoba sydneyensis (Pocock) 
Figures 1-11 
Geophilus sydneyensis Pocock, 1891: 219; Attems 
1914: 133; Attems, 1929: 326. 
Necrophloeophagus sydneyensis (Pocock): Pocock, 
1901: 461. 
Geophilus (Aporophilus) sydneyensis (Pocock): 
Attems, 1903: 262. 
Geophilus hartmeyeri Attems, 1911: 158. figs 11, 12 
(in part); Attems, 1914:127 (in part) Attems 
1914: 133; Chamberlin, 1920: 53 (in part) 
(synonymised by Crabill, 1968: 345). 
Tuoba curticeps Chamberlin, 1920: 35; Attems, 1929: 
338. New synonymy. 
Geophilus (?) sydneyensis Pocock: Chamberlin, 1920: 
54. 
Nesogeophilus hartmeyeri (Attems): Attems, 1929: 
186 (in part); Attems, 1947:118 (in part). 
Algerophilus leplochilus Brolemann, 1931: 311. New 
synonymy. 
Algerophilus hartmeyeri Brolemann, 1931: 314, by 
implication. 
R.E. Jones 
Nesogeophilus leptochilus (Brolemann): Attems, 1947: 
118; Demange 1963: 87. 
Material Examined 
Syntype of Geophilus sydneyensis Pocock 
1 6, Inner Double Bay, Port Jackson, New South 
Wales, Australia (BMNH, accession no. BM 
1879.5.20.13). 
Types of Geophilus hartmeyeri Attems 
Lectotype, Denham, Shark Bay, Western 
Australia, Australia (designated by R. Crabill, 12 
May 1960) (ZIMH). 
1 paratype (A), same data (designated by R. 
Crabill, 12 May 1960) (ZIMH). 
1 syntype, Denham, Shark Bay, Western 
Australia, Australia (NMW, 2006). 
Paratype of Tuoba curticeps Chamberlin 
1 2 Wainoni Bay, Solomon Islands, W.M. Mann 
(MCZ TC-138(2167) and slide 2167a). The holotype 
of Tuoba curticeps has been lost. 
Types of Algerophilus leptochilus (Brolemann) 
The types of this species could not be found in 
the Museum National d'Histoire Naturelle, Paris. 
However I was able to examine samples of 
Nesogeophilus leptochilus which Demange used for 
his 1963 paper. 
Other Material 
Australia: Western Australia: 13 6,8 2, Eagle 
Bluff, 26°06'S, 113°35'E, 25-29 November 1991, 
seagrass litter from littoral zone, M.S. Harvey and 
M.E. Blosfelds (WAM 92/1050-71); 3 6, 2 2, Eagle 
Bluff, 26°06'S, 113“35'E, 25-29 November 1991, 
under rocks, littoral zone, M.S. Harvey and 
M.E.Blosfelds (WAM 92/1072-6); 2 2, Mandu 
Mandu Beach, Cape Range National Park, 29 
February 1986, on beach, under crab exuviae, J.M. 
Waldock (WAM 92/1112-3); 2 6, 1 2, Rosemary 
Island, 6 August 1985, R.P. McMillan (WAM 92/ 
1386-8). 
Papua New Guinea: New Britain: 5 6, 2 2, 
Hoskins, 23 May 1997, on sandy beach under small 
logs, R.E. Jones (R.E. Jones collection). 
Description 
Maximum length 32 mm. 
Number of leg-bearing segments 39-55 (43-49 <J, 
43-55 2 in Australia, 39—41 6 , 41-43 2 in New 
Caledonia, 45 6, 45—47 2 in New Britain and 47 6, 
49 2 in the Solomon Islands). The numbers of legs 
for males and females in this and the following 
species will no doubt be extended as more are 
collected. 
Colour orange yellow throughout. Antennae: 
about 5 times the length of the head capsule, first 
segment as broad as long then from the second to 
the penultimate about twice as long as broad, last 
segment greatly exceeded by the previous two 
(Figure 5). Setae moderate throughout. Head 
