Australasian Tuoba 
337 
capsule as long as broad, the sides being convex 
(Figure 2), Clypeus 1+1 postantennary setae and 
6+6 midlabral setae, there is a slight area of 
reduced areolation on either side in front of the 
labrum. Labrum tripartite, 7-9 teeth on the 
midpiece and 2+2 on the sidepieces (Figure 6). First 
maxilla 3+3 setae on the teleopodites and 4+4 setae 
on the coxal projections, (in both cases there is 1 
large seta and the rest are very small), there is a 
small lappet on the teleopodite. Second maxilla 
bears a simple claw surrounded by a whorl of six 
or seven setae (Figure 4). Forcipules (prehensors / 
poison claws) with concavity of claw smooth and a 
small basal node, reaching almost to the front of 
the head capsule when closed. Chitin lines 
complete or nearly so (Figures 1, 7). Sternites: pores 
throughout forming a ventral band on the sternite 
which becomes two separate groups by the end of 
the animal, also on the procoxa and metacoxa plus 
a few scattered across the front of the anterior 
segments. Areolation less than in T. laticeps 
forming a band at the dorsal and later the ventral 
areas of the sternite. Carpophagus structure 
present until tine transition, the fossa being over 
half as wide as the sternite at its full extent (Figure 
11). Metatergite trapeziform, wider them long (4:3), 
sides slightly convex (Figure 10), metasternite 
trapeziform, the sides being slightly concave. 
Coxopleuron contains about 20-35 glands in a 
rosette arranged around a pit. Last legs of female 
two times the length of the penultimate legs, femur 
2:1 length to breath (Figure 9), last legs of male 
fatter, femur about 1.5 : 1 length to breath (Figure 
8). Anal pores present. Claws present on last legs. 
Remarks 
Brolemann's excellent drawings and the 
specimens of Demangc that I have seen show that 
it is clearly a species of Tuoba. Nesogeophilus 
leptochilus has a lower number of leg pairs than 
the Tuoba sydneyensis of Australia (39—43 pairs) but 
I can find no other differences between them and 
am inclined to amalgamate them as one species, 
T. sydneyensis. Likewise T. curticeps, of which the 
type is lost but a paratype remains, shows the 
characters associated with T. sydneyensis and I 
have also amalgamated them as T. sydneyensis. 
The specimens collected from New Britain also 
appear to be T. sydneyensis with 45-47 pairs of 
legs. The only difference being the pretarsal 
anterior parunguis which is as long as the claw in 
one specimen and may represent the unworn 
state. 
Tuoba laticeps (Pocock) 
Figures 12-23 
Geophilus (?) laticeps Pocock, 1891: 220, pi. 12, fig. 6, 
6a; Chamberlin, 1920: 54. 
Necrophloeophagus laticeps (Pocock): Pocock, 1901: 
463. 
Geophilus (Aporophilus) laticeps (Pocock): Attems, 
1903: 261. 
Geophilus hartmeyeri Attems, 1911: 158. figs 11, 12 
(in part); Attems, 1914: 127 (in part); Attems 
1914: 130; Chamberlin, 1920: 53 (in part) 
(synonymised by Crabill, 1968: 345). 
Geophilus laticeps Pocock: Attems 1914: 130; Attems, 
1929: 328. 
Nesogeophilus hartmeyeri (Attems): Attems, 1929: 
186 (in part). (Attems): Attems, 1947: 118 (in 
part). 
Algerophilus hartmeyeri Brolemann: 1931 by 
implication. 
Nesogeophilus laticeps (Pocock): Crabill, 1962: 506; 
Mesibov, 1986: 54. 
Tuoba laticeps (Pocock): Crabill 1968: 345. 
[Tuoba xylophagus: Crabill, 1968, not (Attems) 1903 
(incorrect synonymy by Crabill, 1968: 345).] 
Material Examined 
Holotype of Geophilus laticeps Pocock 
2, King Island, Bass Strait, Australia, A. Dendy 
(BMNH, accession no. BM 1888.144). 
Syntype of Geophilus hartmeyeri Attems 
1, Albany, Western Australia, Australia (NMW, 
2007). 
Other Material 
Australia: Tasmania: 1 2, Midway Point, S.E. 
Tas., 4 April 1973, under rocks at high tide level on 
shore, Mr G. Prestedge [det. Tuoba sp. Dr R.E. 
Crabtree (sic), 1984] [TM J3139/2, slide prepared 
by R.E. Crabill (labelled 74.11.7/GC-6)]; 1 d, 10 2, 
Midway Point, S.E. Tas., 4 November 1973, under 
rocks on sand, at high water mark, under a 
causeway (= the second causeway), Mr G. 
Prestedge [det. Tuoba sp. Dr R.E. Crabtree (sic), 
1984] (TM J3140); 2 d, 1 2, Brick Maker's Beach, 
Municipality of Circular Head, 21 January 1995, 
under stones on fine shell gravel, rocky shore, 
upper barnacle zone and black obtuse "Littorina", 
S.J. Lewis (QVM 23:17732); 6 d, 5 2, E. end of Bay 
just West of Penguin, 30 January 1995, under 
stones, strand line by rocky outcrop S.J. Lewis 
(QVM 23:17730); 6 d, 5 2, N. end of Ralphs Bay, 
Lauderdale, Hobart, 25 January 1995, under stones, 
strandline, stony beach, S.J. and J.G.E. Lewis (QVM 
23:17733); 1 8, N. corner Prosser Br. Bay, Orford, 
26 January 1995, in small cave 1 x 1 d, under rock 
+ woodlice, crabs, sand hoppers, J.G.E. Lewis 
(QVM 23:17735); 2 d, 2 2, 2 d, 2 2, Swansea, S. 
most part Nine Mile Beach by small rocky prom., 
27 January 1995, under stones, drift zone with 
woodlice, sandhoppers, S.J. Lewis (QVM 23:17736). 
