408 
J.H. Bradbury, W.D. William, 
medial setule. Telson (Figure 3) shorter than broad 
(45:63), sub-equal in length to urosomite 3, cleft 
100%, laterally tumid, the apices rounded or 
slightly notched, bearing a moderately long single 
naked seta and 2 small penicillate setules, and on 
the left lobe a single sub-marginal medial seta at 
M0.7. Lateral penicillate setules absent. 
Description of other material 
Male 'c', 3.5 mm. A1 flagellum of 16 articles, 
aesthetascs present on articles 11, 14, 16; relative 
lengths of peduncular articles; 19:21:6 (compares 
with 24:27:10, converts to 2 4:27:8), length of 
flagellum relative to peduncle; 77:46 (compares 
with 115:61, converts to 115:68), and accessory 
flagellum reaching to end of article 1 of primary 
flagellum. 
Distribution 
Bundera Cenote (22°25’S, 113°46'E), North West 
Cape, Western Australia. 
Relationship 
Liagoceradocus branchialis, like other species 
assigned to the genus, differs from Barnard's (1965) 
original definition in minor ways. Rudimentary 
inner lobes are present on the lower lip, palmar 
setae are found on the second gnathopod, and 
there are 5 rather than 6 D setae on the third article 
of the mandibular palp, and which article is not 
longer than article 2. In keeping with the original 
definition of the genus the telson is deeply cleft, 
although, unlike in Barnard’s revised (1977) 
concept, lateral setae (spines) are absent. L. 
branchialis differs also from L. pusillus Barnard in 
that the apex of the third maxillipedal palp article 
is not apically produced, the rostrum is reduced, 
but not entirely absent; the third epimeron is not 
acuminate; there are no dorsal spines on 
urosomites 1-2; sparse setae on the inner face of 
the inner plate of the second maxilla; post-ventral 
lobes on the second article of the fifth and sixth 
pereopods. 
L. branchialis differs from L. lonomaka Barnard in 
the presence of a small rostrum; relatively shorter 
antennae; symmetric molarial setae; the third 
article of the maxillipedal palp is similarly armed 
but not as broad; the uropods are less setate, the 
peduncular spines of U2 are simple not forming a 
comb; the telson is without medial and lateral 
spines; the cuticle smooth, with few setae; the 
second article of P5 is broader than on P4. 
L. branchialis differs from L. acutus Andres in the 
length of the A1 accessory flagellum which is 
relatively shorter; A2 is composed of peduncle and 
6 rather than 14 flagellar articles, the gland cone is 
less tapered; the upper lip has less apical setation; 
mandibular palp article 3 has fewer D setae; the 
inner lobe of the first maxilliped has fewer plumose 
setae; maxillipedal palp article 3 is not produced 
apically, and spines of the plates are naked, not 
setulate or rastellate; coxal plates bear fewer seta»>; 
G1 is without palmar comer spines, the palmar 
margin is without spinules or bifid spines; the legs 
are less setose or spinous; epimera are without ah 
oblique ridge, the post ventral corners without 
teeth or acumination, ventral margins are without 
spines or setae; peduncles of pleopods bear 2 
retinacula each without accessories, setae are all 
simple and plumose; basofacial spine of U1 is 
weak; peduncles and rami of uropods with fewer 
spines and setae; the telson is without lateral 
spines. 
L. branchialis differs from L. dentiferus Ledoyer ih 
that the second article of P5 is weakly lobate; the 
width of C4 is less than its depth; the palm of the 
second gnathopod has fewer spines or setats; 
telsonic lobes are without lateral or medial spines. 
The two are similar in that the third article of the 
maxillipedal palp is curved, and the dactyl bears 
accessory spinules. 
L. branchialis differs from L. unciferus Stock and 
Iliffe in possessing a shorter A1 accessory 
flagellum, aesthetascs longer than flagellar 
segments; the gland cone of A2 is less pointed; 
fewer setae on articles 2-3 of the mandibular palp; 
less apical setae on the palp of the first maxilla; the 
G1 is without spines at the palmar corner, and 
without bifid spines; G2 dactyl bears larger 
recumbent inner tooth spines, there are fewer 
palmar spines, the palmar comer bears bifid rather 
than naked spines, as well as long 'hadziid' setae; 
coxae and Pereopods bear fewer marginal setae, 
coxa 4 is wider than long; epimeral plates 1-2 are 
post-ventrally rounded rather than pointed, 
without ventral setae or spines; pleopods are 
unmodified, similar, without bifid setae or 
accessory retinacula; uropods are less spinous; 
telson is without medial or lateral spines, with 
apical penicillate setules. 
L. branchialis differs from L. lobiferus Stock and 
Iliffe; the body is similarly armed except for the 
absence of dorsolateral spines on urosomites 1-2; 
appendages are less spinous or setate; U3 is 
relatively longer; A1 peduncle article 2 is longer 
than article 1, the accessory flagellum is shorter 
than the first article of the primary flagellum, the 
primary flagellum has more articles but the same 
relative length, the aesthetascs are longer; A2 gland 
cone is broader and less pointed apically, the 
flagellum of similar numbers of articles; 
mandibular molar well developed, palps with 
fewer setae; first maxilla inner lobe with more 
plumose setae although similarly arranged, the 
processes of the spines of the outer plate are 
similar, the palps with fewer terminal spines; 
second maxilla bearing more setae both plumose 
