68 
copepods, melitid amphipods (undescribed genus; 
W. D. Williams, pers. comm.), Halosbaena tulki 
Poore and Humphreys (Thermosbenacea), 
Stygiocaris lancifera Holthuis (Decapoda: Natantia), 
Hydracarina (Acarina), Milyeringa veritas Whitely 
(Perciformes: Eleotridae) and Ophistemoti candidum 
(Mees) (Synbranchiformes: Synbranchidae). 
Mesocyclops brooksi is known only from the type 
locality, Bobs Well, a hand-dug well in the 
Ashburton River valley, a river now remote from 
Cape Range but which formerly flowed past the tip 
of the peninsula (Wyrwoll et al. 1993). The water 
temperature was 25.0°C and the salinity was 1121 
mg LA 
Apocyclops dengizicus was found only in a small 
piezometric tube located in a recently dug test pit 
in a back dune area adjacent to the ocean. The 
water temperature was 26.9°C and the salinity was 
22,000 mg LA 
Microcyclops mricans was collected in a variety of 
habitats containing waters ranging in temperatures 
between 23.9 and 26.9°C and salinity between 99 
and 9448 mg LA These sites included a bore and a 
hand-dug well at the base of the Cape Range 
peninsula, a hand-dug well on the Ashburton River 
and, interestingly, water in the dark zone of an 80 
m deep cave (C-18) within central Cape Range- 
where it occurs with a second species of the 
undescribed genus of melitid amphipod mentioned 
above, the only other aquatic species known from 
the range. This cave contains a diverse troglobitic 
fauna derived from a rain forest (both temperate 
and tropical) litter fauna - the area is now arid 
(Humphreys 1993a, 1993c). In addition, the aquatic 
insect Copelatus irregularis Macl. (Coleoptera: 
Dytiscidae) has been recorded in this cave (itself a 
big range extension from the Kimberley, central 
Australian Ranges and North-east Queensland). 
MATERIAL AND METHODS 
Groundwater was sampled for fauna at 261 sites 
on the Cape Range peninsula and in the Ashburton 
River valley in 1993; 185 of these sites yielded 
stygofauna (Figure 29). The groundwater was 
accessed through bores, wells, soaks, piezometers 
and caves. Bores were sampled for fauna using a 
plankton net with a 125 pm mesh and of a size 
suitable for the bore, the nets ranged from 30 mm 
to 180 mm in diameter. Wells were sampled by 
hand nets (125 pm mesh) and by Cvetkov (1968) 
phreatobiological nets (300 mm diameter, 250 pm 
mesh). Samples were variously taken through the 
entire water column and from various depths 
within the water column — occasionally the outflow 
from bore pumps was sampled by straining 
through a net (125 pm mesh). In all situations 
baited traps were sometimes used. 
Comprehensive water analyses are available for 
G.L. Pesce, P. De Laurentiis, W.F. Humphreys 
seven of the eleven sites mentioned in this paper 
(Humphreys 1994). Where direct measurements 
were not made the salinity (total soluble salt: TSS) 
was estimated from the electrical conductivity 
measured in the field. 
Permanent mounts were made in commercial 
polyvinyl-lactophenol medium. Dissected 
specimens were drawn at magnification of 400x 
and lOOOx, the latter using an oil immersion lens 
and "camera lucida" mounted on a Leitz Laborlux 
D phase-contrast microscope. Type material is 
deposited in the Western Australian Museum 
(WAM). 
Terminology applied to body and appendages 
according to Huys and Boxshall (1991). 
SYSTEMATICS 
Family Cyclopidae Burmeister, 1834 
Genus Metacyclops Kiefer, 1927; sensu Lindberg, 
1961 
Metacyclops mortoni sp. nov. 
(Figures 1-15) 
Material Examined 
Holotype 
$ (WAM 183-94), Kudamurra Well (C-25), Cape 
Range peninsula. Western Australia (station BES 
2199), Australia, 21°54'S, 113°49'E, 1 June 1993, R.D. 
Brooks. 
Paratypes 
Australia: Western Australia: 1 <J (allotype) 
(WAM 2-184), 2 9 (WAM 185, 186-94), Geebera 
Well, Ashburton River (BES 2321), 22°06'S, 
115°08'E, 26 June 1993, R.D. Brooks and W.F. 
Humphreys; 2 2, (WAM 187, 188-94), Five Mile 
Well (C-273) (BES 2196), 21°51'S, 113°52'E, 1 June 
1993, R.D. Brooks; 2 2 (WAM 189, 190-94), Tulki 
Well (C—149) (BES 2213), 22°05S, 113°54'E, 2 June 
1993, R.D. Brooks; 2 2 (WAM 191, 192-94), 
Waroora Well, Pt. Maud (BES 2227), 23°09'S, 
113°48'E, 11 June 1993, R.D. Brooks. 
Description 
Female 
Length of holotype, excluding antennule and 
caudal setae, 555 pm; range of length of 6 
paratypes 540-560 pm. Body widest at first 
prosomite. Genital segment about as long as broad, 
anterior 2/3 faintly expanded laterally; seminal 
receptacle with well developed subcircular anterior 
expansion, lateral canals almost horizontal. First 
urosomites naked, anal somite with small spinules 
along posterior margin; anal operculum not well 
defined. 
