126 
M.R. Yardin, B.J. Richardson 
W.A. Our thanks go to George Kendrick (Western 
Australia Museum), Winston Ponder and Ian Loch 
(The Australian Museum, Sydney) for useful 
literature and comments. We are also grateful to 
Elizabeth Turner (Tasmanian Museum and Art 
Gallery) and Ben McHenry and Neville Sledge 
(South Australian Museum) for information on the 
Museum records. This research would not have 
been successful without the expert guidance of 
Patricia Dixon (Centre for Marine Science, 
University of New South Wales) on the ecological 
genetics of A. trapezia. Her help is gratefully 
acknowledged. 
REFERENCES 
Avise, J.C. (1994). Molecular Markers, Natural History and 
Evolution. Chapman and Hall, Inc., New York. 
Ayala, F.J., Tracey, M.L., Hedgecock, D. and Richmond, 
R.C. (1975). Genetic differentiation during the 
speciation process in Drosophila. Evolution 28: 576- 
592. 
Barton, N.H. and Charlesworth, B. (1984). Genetic 
revolutions, founder effects, and speciation. Annual 
Review in Ecology and Systematics 15: 133-164. 
Beu, A.G. and Maxwell, P.A. (1990). Cenozoic Mollusca 
of New Zealand. New Zealand Geological Survey 
Paleontological Bulletin 58: 518p. 
Bryner, V.F., and Grant-Mackie, J.A. (1993). The bivalve 
Anadara trapezia and other macrofossils in tuffs of 
Motukorea volcano, Auckland. Tane 34: 123-131. 
Burton, R.S. (1986). Evolutionary consequences of 
restricted gene flow among natural populations of 
the copepod, Tigriopus califomicus. Bulletin of Marine 
Science 39: 526-535. 
Carson, H.L. (1975). The genetics of speciation at the 
diploid level. American Naturalist 109: 83-92. 
Cotton, B.C. (1957). Records of uncommon Southern 
Australian molluscs. Records of the South Australian 
Museum 13: 117-130. 
Cotton, B.C. (1961). South Australian Molluscs: Pelecypoda. 
Government Printer, Adelaide. 
Crozier, M. (1962). New Zealand occurrences of the 
Sydney Mud Cockle, Anadara trapezia (Deshayes). 
Records of the Dominium Museum 4: 143-148. 
Dixon, P.I. (1975). The ecological genetics of Anadara 
trapezia with particular reference to its haemoglobin. 
PhD Thesis. University of New South Wales. 
Dobzhansky, T. (1951). Genetics and the Origin of Species. 
3rd ed. Columbia University Press, New York, 
Eldredge, N. and Gould, S.J. (1972). Punctuated 
equilibria: an alternative to phyletic gradualism. Pp. 
82-115. In T.J.M. Schopf (ed.). Models in Paleobiology. 
Freeman, Cooper and Co., San Francisco, CA. 
Goede, A., Murray-Wallace, C.V. and Turner, E. (1993). 
A diverse Holocene molluscan fauna, including 
Anadara trapezia, from Royal Park, Launceston, 
Tasmania. Papers and Proceedings of the Royal Society 
of Tasmania 127: 17-22. 
Gosling, E.M. (1994). Speciation and species concepts in 
the marine environment. In A.R. Beaumont (ed). 
Genetics and Evolution of Aquatic Organisms: I -15 
Chapman and Hall, London. 
Gould, S.J. and N. Eldredge, N. (1977). Punctuated 
equilibria: the tempo and mode of evolution 
reconsidered. Paleobiology 3: 115-151. 
Grant, V. (1963). The Origin of Adaptations. Columbia 
University Press, New York. 
Grant-Mackie, J.A. and Cook, S.De C. (1990). A late 
Quaternary Anadara —bearing deposit disturbed by 
Rangitoto lava. New Zealand Natural Sciences 17: 73- 
79. 
Heatwole, H. (1987). Major components and 
distributions of the terrestrial fauna. Pp.101-135. In 
Dyne, G.R. and Walton, D.W. (eds). Fauna of 
Australia. General Articles, Vol. 1 A. Bureau of Flora 
and Fauna, Canberra. Australian Government 
Publishing Service, Canberra. 
Hedgecock, D. (1986). Is gene flow from pelagic larval 
dispersal important in theadaptation and evolution 
of marine invertebrates? Bulletin of Marine Science 39: 
550-564. 
Huxley, J.S. (1942). Evolution, the Modern Synthesis. R ev . 
Ed. Harper, New York. 
Johnson, M.S. and Black, R. (1990). Genetic divergence 
of venerid clams in Shark Bay, Western Australia. 
Pp. 159-168 In P.F. Berry, S.D. Bradshaw, and B.R. 
Wilson (eds). Research in Shark Bay. Report of the 
Franee-Australe Bicentenary Expedition Committee 
Western Australian Museum. 
Kendrick, G. (1990). A Pleistocene molluscan fauna with 
Anadara trapezia (Deshayes) (Bivalvia: Arcoida) from 
the Dampier Limestone of Shark Bay, Western 
Australia. Pp. 33-48 In P.F. Berry, S.D. Bradshaw, 
and B.R. Wilson (eds). Research in Shark Bay. Report of 
the France-Australe Bicentenary Expedition Committee. 
Western Australian Museum. 
Kendrick, G. and Wilson B.R. (1959). Anadara trapezia 
(Mollusca - Pelecypoda) found living in South- 
Western Australia. The Western Australian Naturalist 
6 : 191-192. 
Kimura, M. and Ohta, T. (1973). Mutation and evolution 
at the molecular level. Genetics 73 (suppl.): 19-35 
Ludbrook, N.H. (1984). Quaternary molluscs of South 
Australia. South Australian Depl.of Mines and Energy 
Handbook No. 9. Adelaide, Government Printer. 
Macpherson, J.H. (1966). Port Phillip Survey 1957-1963. 
Mollusca. Memoirs of the National Museum of Victoria 
27: 201-263 
Macpherson, J.H. and C.J. Gabriel (1962). Marine 
Molluscs of Victoria. Melbourne University Press. 
Manchenko, G. (1994). A Handbook of Detection of 
Enzymes on Electrophoretic Gels. CRC Press. Ann 
Arbor, London. 
Mayr, E. (1942). Systematics and the Origin of Species. 
Columbia University Press, New York. 
Mayr, E. (1954). Change of genetic environment and 
evolution. Pp. 157-180 In J. Huxley, A.C. Hardy, and 
E.B. Ford (eds). Evolution as a Process: Allen and 
Unwin, London. 
Mayr, E. (1963). Animal Species and Evolution. Harvard 
Univ. Press, Cambridge, MA. 
