A. Wakefield 
Novapex 13(2): 69-74, 10 juin 2012 
between the two main ones. Most specimens of M 
bicatenata are dead collected and worn - it may be 
that fresh specimens hâve traces of extra fine spiral 
Unes in the surface layers of the shell. These markings 
are not as defined or as bold as those found in M. 
lemaitrei. 
M. bicatenata is very distinctive and is unlikely to be 
confused with anything else. M. nevillana Kilbum, 
1977 (7.6 x 4.5 mm) as yet only known from its 
holotype from the eastem Cape, is much smaller than 
M. bicatenata , has eight rows of large charcoal grey 
spots on a white background, a much thicker shell and 
because of its denticulate lip is better placed in the 
genus Glabella Swainson, 1840 (in the G. obtusa 
Sowerby, 1846 complex). 
Marginella lemaitrei Liltved & Millard, 1994 
Figs 22-26 
Marginella lemaitrei Liltved & Millard, 1994: 3, 4, fig 
1 . 
Type material. 1 ad. sh., holotype, 18.4 x 10.0 mm, 
ex. pisce , trawled approx 100m, Cape St. Blaize, 
Southern Cape, South Africa, SAM A3 7572 (Figs 22- 
24) ; 1 ad sh., paratype, 17.1 x 8.9 mm, ex. pisce , 
trawled approx 100m, Cape St. Blaize, Southern Cape, 
South Africa, NM E7213/T 184. 
Other material examined. 1 ad. sh., unmeasured, no 
data, photographed by Markus Lussi (Figs 25, 26). 
Specimen figured on p. 6, 7, fig. Db, in Millard, 1981 
(21x11mm), ‘ ex . pisce ’ off the Cape. 
Specimen image by Brian Hayes (2011). 
Type locality. Cape St Blaize, Southern Cape, South 
Africa. 
Distribution. Restricted to the Agulhas Bank off the 
Southern Cape, South Africa. 
Descriptive notes. Shell length 17.1 — 21.0 mm, W:L 
ratio 52-54% (mean 53%), straw coloured to greyish, 
thin, smooth, satin, elongate-biconic, of 4.5 whorls 
including paucispiral protoconch, weakly shouldered, 
tapering to a narrow anterior end. Suture not 
impressed. Lip with smooth, evenly thickened, white 
margin, smooth internally, with extemal varix groove. 
Plications strong, evenly spaced plications filling over 
half of aperture. First two oblique, third and fourth 
straighter. Columella slightly concave. Aperture 
narrowing posteriorly, otherwise of even width. 4-8 
spiral rows of fine grey spots and streaks on body 
whorl. Holotype has 8 rows; 4 anteriorly, 2 at mid- 
body, 2 on the spire. Paratype has 4 rows with less 
frequent, more blurred markings. Axial pattern of pale 
grey narrow to wide flammules on body whorl. 
Remarks. According to Liltved and Millard (1994: 4), 
M. lemaitrei occurs sympatrically with M. bicatenata 
(‘M tomlini form’) off the Southern Cape. In fact the 
types of M lemaitrei and the holotype of M. tomlini 
are syntopic. The types of M. lemaitrei were taken 
from the stomach contents of the fish Congiopodus 
tonms Walbaum. 
M. lemaitrei differs from the ‘M. tomlini form’ of M 
bicatenata in that it is generally slightly larger, more 
slender (less shouldered), has multiple rows of dashes 
and dots rather than two, and often has an additional 
axial pattern of pale grey narrow to wide flammules. 
The slender thin shell and axial flammules are also 
characters seen in Marginella diadochus Adams & 
Reeve, 1848 and therefore the possibility of an 
ancestral Iink of the M bicatenata group to the M. 
mus ica group should be considered. 
Marginella seccombei n. sp. 
Figs 28-39 
Type material. 5 ad. sh., dredged, preserved dry, off 
Central KwaZulu-Natal in 150m; holotype, 12.61 x 
7.46 mm, NM W8641/T2985 (Figs 27-30); paratype 
1, 13.29 x 8.37 mm, NM, W8642/T2986, (Figs 31, 
32); paratype 2, 12.48 x 8.28 mm, AWC (Figs 33, 34); 
paratype 3, 10.88 x 6.35 mm, TMC (Figs 35, 36); 
paratype 4, 13.75 x 8.48 mm, 
NMW.Z.2012.016.00001 
Type locality. Off central KwaZulu-Natal, South 
Africa. 
Other material. 1 ad. sh. unmeasured, off Durban, 
KwaZulu-Natal, photographed by Markus Lussi (Figs 
37, 38) 
Distribution. Off Central KwaZulu-Natal in 100-150 
métrés. 
Description. Shell moderately sized (L=12.6 mm), 
W:L ratio 58 - 66 % (mean 61.4%), solid, biconic, of 
4 Vi whorls including paucispiral protoconch, 
posteriorly tapering to moderately wide columella 
base. Pale straw coloured with two spiral bands of 
very short axial ly oriented red-brown dashes 
encircling body whorl. Anterior row appears darker 
due to dashes being doser together. Dashes regularly 
spaced and shaped ventrally, becoming more 
separated and irregular from mid-dorsum to lip. 
Anterior (mid-body) row of 24 closely spaced dashes 
emerges immediately posterior to 4 th plication. 
Posterior row of 20 more spaced marks at shoulder on 
body whorl, continuing for another 2 % whorls onto 
spire. Lip, base of columella and plications white. 
Shoulder rounded, spire moderately elevated, suture 
présent, spire whorls shouldered. Posterior end of lip 
inserts at shoulder at level of posterior spiral band. 
Columella slightly concave anteriorly, pariétal surface 
very slightly convex. Aperture widest in middle third, 
twice as wide as the lip, narrowing posteriorly and 
anteriorly to as wide as the lip. Lip evenly convex in 
71 
