A. Wakefield 
A review of Marginella bicatenata 
However, it lacks the two main spiral bands of brown 
axial markings and it has a weakly denticulate lip. 
Although smaller than M. seccombei, M monicae is a 
shell of comparable shape though it is thicker and 
slightly pyriform. Although mainly lacking a colour 
pattern, it has a faint trace of dark markings (not clear 
spots or axial streaks) at the shoulder and at the 
anterior end in the same place as the spiral rows in M 
seccombei. It appears to be the species closest 
morphologically to M. seccombei discovered to date. 
M tuguriana Lussi,1993, another benthic species 
occurring in northern Natal and possibly ranging 
further north into Mozambique, also has a faint 
anterior band, but its extremely biconic morphology, 
thicker shell and labial markings distinguish it clearly 
from M. seccombei n. sp. 
Acknowledgements 
Dr. Igor Muratov (KwaZulu-Natal Muséum) kindly 
provided the images of M bicatenata (Figs 3-12). 
Linda Davis (Collections Manager, KwaZulu-Natal 
Muséum), Liz Hoenson (Iziko, South African 
Muséum), and Harriet Wood (Amgueddfu Cymru, 
National Muséum of Wales) generously provided 
access to type material and/or information. 1 would 
like to express spécial thanks to Alwyn Marais (Centre 
for Molluscan Studies, South Africa) for the images of 
the holotype of M tomlini (Figs 13-15), and to 
Markus Lussi (Durban) for his opinions on the 
distribution of M. bicatenata, for his excellent images 
(Figs 16-21, 25, 26, 37 and 38) and for being my 
referee. A final thanks to Alan Seccombe (Cape 
Town) for providing the author with type material ot 
M seccombei. 
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Liltved, W.R., & Millard, V.G. 1994. Five New 
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Millard, V. G., 1981. Marginellidae of South Africa. 
The Strandloper , 206: 1-12, 2 pl. 
Shackleford, L.J., 1916. Two New Species of 
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