B. M. Landau & L. T. Groves 
Cypraeidae from the early Miocene of northem Venezuela 
Dimensions and type material. Holotype; LACMIP 
12431, length 42.1 mm (Figs 23-24); paratype I 
ANSP 315087, length, 42.0 mm (Figs 21-22); 
paratype 2 USNM 562582, length, 38.5 mm, paratype 
3 NHMW 2010/0036/0001 (ex. BL coll.) length 43.4 
mm (figs. 25-28). 
Other Material. Panama, three incomplète specimens 
PPP 02168 (NMB 18667); three incomplète 
specimens PPP 02119 (NMB 17637); two specimens 
PPP 00490 (NMB 17871); four specimens PPP 00487 
(NMB 17868); one specimen PPP 1080 (NMB 
18261); one specimen PPP 00224 (NMB 17642); two 
specimens PPP 01034 (NMB 18258); one incomplète 
specimen PPP 01078 (NMB 18325) ail NMB coll.; 
one specimen locality ANSP 1731 (internai mould); 
one specimen locality LACMIP 16936; five 
specimens locality LACMIP 17908 (= TU 757); five 
specimens locality LACMIP 17909 (= TU 958); two 
specimens locality LACMIP 17910 (= TU 960); two 
specimens locality LACMIP 17911 (= TU 1432); one 
specimen from LACMIP 17912 (= TU 1433); Cativa, 
Colon, BL coll. (Ail Gatun Formation); Colombia, one 
specimen locality UCMP 154008 (fragment), Narino 
Dept.; Ecuador, one incomplète specimen PPP 03391 
(NMB 19122); one specimen AGC 99-102, NMB 
coll.; Baja California Sur, Mexico, one specimen 
locality USNM 1/1238 (ex USGS), San Ignacio 
Formation!?). 
Type locality. Angostura Formation (late Miocene), 
Loc. LACMIP 16943 [= TU 1507], just east of Rio 
Verde, approximately 30 km east of Rio Esmeraldas 
Esmeraldas Province, Ecuador. 
Type stratum. Angostura Formation (late Miocene). 
Description. Shell medium-sized for genus, solid, 
triangular-shaped, dorsum very strongly humped 2 / 3 
distance from the anterior end, bearing two groups of 
small warty tubercles, left group larger and slightly 
anterior to right; spike-like projection at centre of apex 
rises 2 mm above dorsal surface; spire depressed, 
covered by adapical callus; sides rounded, strongly 
callused, with the callus ascending progressively 
adapically to just below the apex posterior to the 
dorsal hump, callus edge poorly delimited; ventrum 
flattened, slightly convex in profile; aperture almost 
straight, anterior third weakly dilated; anterior canal 
deep, abaxially asymmetrical and abapically 
orthogonally truncated, flanked on either side by 
flattened spatulate horizontal expansions produced 
from the abapical tips of the inner and outer lips; 
posterior channel deeper, limited by parallel lips; 
about 4 mm width weakly concave channel between 
terminal ridge and first columellar tooth, followed by 
14 stout, becoming weak posteriorly, short columellar 
teeth, extending a short distance onto the ventrum and 
into the aperture; terminal ridge obsolète, merged into 
smooth abapical edge of fossula; fossula small, 
concave, smooth; labral teeth heavy, 14 in number, 
evenly distributed, becoming weak posteriorly. 
Ventral and dorsal zones striped, marginal callus 
lighter coloured, dorsum with irregular stripes of dull 
red-orange and tan. 
Shell Formula. 45.3 (80.0-60.0) 21; 14 
specimen 
collection number 
length 
width 
Height 
LT 
CT 
holotype (Figs 23-24) 
LACMIP 12431 
43.5 
34.5 
27.1 
18 
# 
paratype 1 (Figs 21-22) 
ANSP 315087 
42.1 
34.1 
24.6 
14 
13 
paratype 2 
USNM 562582 
46.0 
36.9 
24.3 
# 
# 
paratype 3 (Figs 25-28) 
NHMW 2010/0036/0001 
43.4 
31.1 
25.6 
18 
13 
CAS 66589.06 
37.5 
29.7 
22.6 
# 
# 
USNM 562581 
52.7 
44.0 
31.8 
# 
# 
BL coll. 1 
38.4 
31.6 
21.4 
18 
13 
BL coll. 2 
42.4 
35.1 
23.1 
17 
12 
BL coll. 3 
57.6 
45.1 
31.8 
18 
12 
BL coll. 4 
48.8 
41.0 
28.5 
17 
13 
BL coll. 5 
46.6 
35.5 
26.7 
17 
12 
DF B coll. 
No. 7406-1 
48.5 
39.8 
26.3 
17 
11 
DFB coll. 
No. 7406-2 
38.8 
32.7 
22.1 
16 
12 
Table 2. Dimensions and number ot teeth; Muracypraea woodringi n. sp. 
Discussion. Muracypraea woodringi is described 
the forms noted by Groves (1997) as M. “heneke 
hom the Gatun Formation of Panama, the Angosl 
Formation of Ecuador, and Miocene strata 
southwestern Colombia. A poorly preser 
specimen from Southern Baja California Sur, Mex 
may also represent M. woodringi. Mnracypn 
woodringi is most similar to M. isthm 
(Schilder, 1927) [= Cypraea henekeni Sowb. var 
6 
Brown & Pilsbry (1911:356-357, pi. 26, figs. 9-10)] 
also from the Gatun Formation, which has a broad, 
smooth rounded, poorly delimited dorsal gibbosity 
centrally positioned (Figs 23-24). These shells were 
illustrated by Woodring (1959; pi. 31, figs 6-10). The 
specimen illustrated by Brown & Pilsbry (1911: pi- 
26, figs. 9-10) clearly has a single dorsal gibbosity. 
Untortunately, the holotype is missing. Brown & 
Pilsbry (191 1: 356) noted that the “aperture is like that 
