B. M. Landau & L. T. Groves 
Cypraeidae from the early Miocene of northem Venezuela 
Description. Shell medium-sized for genus, solid, 
subcylindrical to rotund, posteriorly swollen, in latéral 
profile dorsal curvature regularly attenuated 
abapically, spire weakly projecting, covered by 
adapical callus; sides rounded, weakly callused; 
ventrum flattened, slighlly convex in profile; aperture 
parasigmoidal, anterior third dilated by hemispherical 
expansion; siphonal canal deep. abaxially 
asymmetrical and abapically orthogonally truncated; 
exhalant channel deeper, limited by parallel lips; four 
or five anterior columellar teeth, most anterior 
thinnest, strengthening adapically; other numerous 
inner teeth parallel, extending deep within the 
aperture, bituberculate, about 6 mm in length in the 
columellar area, abrupt, extending a short distance 
over the angulation of the basal and columellar planes 
onto the ventrum where they are slightly coarser; 
terminal ridge obsolète, merged into abapical edge, 
fringed with denticles from fossula on inner margin; 
fossula deep, concave, spoon-like, covered by narrow 
ridges, sometimes interrupted, joining fossular 
denticulation to anterior columellar teeth; 22-30 labral 
teeth, anterior third of labral teeth stretched out on 
constricted, depressed, hemicircular area; remainder 
heavier, outer lip bevelled inwards, with teeth 
extending across the bevelled inner portion, 3-4 ram in 
length; ventral and dorsal zones spotted, marginal 
callus lighter coloured, under UV light spire blotch 
présent. 
Shell Formula. 48.7 (63.0 - 50.1 ) 20: 18 
Specimen 
collection number 
length 
width 
height 
LT 
CT 
holotype (Figs 58-60) 
NMB H18402 
53.1 
34.5 
29 
23 
20 
NMB lot 17516 
35.3 
22.4 
18.2 
22 
17 
NMB lot 17516 
35.7 
22 
18.7 
# 
# 
NMB lot 17516 
43 
26.8 
# 
23 
26 
paratype 1 (Figs 61-64) 
NHMW 2010/0036/0008 
48.2 
32.6 
25.7 
23 
23 
paratype 2 (Figs 65-68) 
NHMW 2010/0036/0009 
47.2 
29.7 
23.2 
23 
25 
paratype 3 (Figs 69-70) 
NHMW 2010/0036/0010 
49.7 
30.4 
26.3 
25 
24 
BL coll. 1 
57.9 
36.7 
28.3 
26 
23 
BL coll. 2 
56.1 
34.6 
26.9 
27 
21 
BL coll. 3 
55.8 
35.4 
28.2 
27 
24 
BL coll. 4 
58.9 
36.3 
29.7 
24 
23 
BL coll. 5 
57.4 
39.4 
31.1 
23 
20 
BL coll. 6 
55.9 
34.3 
27.6 
23 
21 
BL coll. 7 
55.6 
36 
29.7 
26 
23 
BL coll. 8 
60.2 
37.5 
29.4 
30 
?8 
BL coll. 1 1 
48.7 
30.9 
25.1 
22 
25 
BL coll. 12 
47.8 
31.8 
24 
23 
20 
BL coll. 13 
45.1 
30.6 
25 3 
23 
20 
BL coll. 14 
44.2 
28.2 
22 
24 
22 
BL coll. 16 
44.8 
27.2 
20.9 
25 
26 
dL coll. 1 7 
51.5 
31.4 
25 2 
?5 
23 
BL coll. 18 
49.5 
32.8 
25 3 
IA 
?1 
BL coll. 19 
40.1 
24.3 
18 9 
BL coll. 20 
37.1 
22 
17 3 
24 
If) 
BL coll. subadult 21 
DFB coll. 
no. 6578 
39.5 
55.1 
25.1 
38.4 
18.8 
30.4 
25 
24 
24 
17 
Table 5. Dimensions and number of teeth; Trôna ingrami n. sp 
Text-figure 3 (opposite). Trôna ingrami n sp 
Paratype 1 NHMW 2010/0036/0008 (NMMW coll.; 
ex. BL coll. no. 9), length 48.2 mm. lower shell bed, 1 
km Southwest of Casa Cantaure, about 10 km west of 
Pueblo Nuevo, Paraguanâ Peninsula, Falcôn State 
Venezuela ’ 
16 
