C. VlLVENS, F. SWINNEN & F. DENIZ GUERRA 
Novapex 12(1-2): 49-55, 10 mars 2011 
Clelandella miliaris (Brocchi, 1814) 
Figs 7-14, Text Figs 1, 3 
Trochus miliaris Brocchi, 1814: 353, pl. 6, fig. 1. 
Type locality: northern Italy, no 
specified locality; lectotype from qualification as 
holotype. 
Trochus clelandi — W. Wood, 1828: 16, pl. 5, fig. 15. 
Trochus millegranus - Philippi, 1836: 183, pl. 10, fig. 
25. 
Calliostoma clelandi - Kaicher, 1979: card#2094. 
Calliostoma miliaris - Poppe & Goto. 1991: 74, pl.6, 
fig. 9. 
Clelandella miliaris - Giannuzzi-Savelli et al., 1994: 
88, figs.279-281. 
Clelandella miliaris - Cretella et al., 1999 : 12-1A, 
figs. 13-15, 17-18, 25,28,33-34. 
Calliostoma miliaris - Ardovini & Cossignani, 2004: 
71. 
Clelandella miliaris - Gofas, 2005: 134-136, figs. 1— 
3, 12A-B, 13A-B. 
Clelandella miliaris - WoRMS, 2011 : 
p=taxdetails&id= 141774. 
Material examined. France. Britanny, Finistère, near 
Ouessant, +/- 100 m, 40 dd. - Roscoff, +/- 3 m, 25 lv. 
Italy. Tuscany Archipelago, 2 lv. Spain. Off Malaga, 
2 lv. West Africa. Off Mauritania, 17°45'N, 16°25'W, 
3 lv, 1 juv lv. 
Distribution. North-eastern Atlantic, from Norway to 
West Africa and Mediterranean Sea, 35-800 m. 
Remarks. The main characteristics of this species 
are : 
- height up to 14 mm, width up to 13.8 mm; 
- a high spire, a conical shape, with up to 7 fiat 
whorls; 
- strongly prosocline axial threads and up to 8 spiral 
cords, granular by intersection with axial threads; on 
First whorl, 4 cords Pl, P2, P3 and P4 appearing 
immediately, smooth; Pl slightly weaker than other 
cords; cords granular by intersection with axial 
threads as soon as second whorl; secondary cords S2 
and S3 appearing by intercalation; P4 thicker on third 
whorl, dividing into 2 cords on fourth whorl, up to 10 
cords on last whorl of big specimens, producing a 
thick rim making carina; 
- columella nearly straight, oblique, with a weak 
médian swelling; 
- base almost fiat, with 8 to 13 granular spiral cords; 
- no umbilicus or very narrow umbilicus reduced to a 
small slit. 
Nicklès (1950) reviewed the known marine species 
of Western and Equatorial Africa but curiously didn't 
mention this species. Kaicher (1979) still used the 
synonym name. At last, Cretella et al. (1999) 
produced an accurate study of this species. 
This is a variable species (Gofas, 2005) regarding size 
and colour, also sometimes with a slightly 
cyrtoconoidal shape instead of a conical one and a 
subangular instead of angular periphery (some 
specimens from western Africa). Gofas mention even 
(without illustrations) some specimens from Ivory 
Coast with a narrow umbilicus. 
Clelandella artilesi n. sp. 
Figs 1-4, Table 1, Text Fig. 2, 3 
Type material. Holotype (6.9 x 7.2 mm) 1RSNB 
1G.31784/MT2330 for database DaRWIN. Paratypes: 
1 MNHN 23351, 2 coll. F. Deniz Guerra, 2 coll. 
F.Swinnen, 1 coll. C.Vilvens. 
Type locality. West Africa. Off Western Sahara, 50- 
60 m. 
Material examined. West Africa. Off Western 
Sahara, 12/1999, 50-60 m, 2 dd sub, 3 dd juv. - Off 
Western Sahara, 7/2001, 50-60 m, 3 dd (with holotype 
and 2 paratypes). - Off Western Sahara, 1993-1999, 
50-70 m, 15 dd, 7 dd sub, 2 dd juv (with 4 paratypes). 
Distribution. West Africa, off Western Sahara, 50-60 
m. 
Diagnosis. A typical cantharid species with an 
elevated, conical spire, up to 6 even granular spiral 
cords on the whorls, an angular periphery, up to 6 or 7 
smooth spiral cords on the base and a very narrow or 
closed umbilicus. 
Description. Shell of moderate size for the genus 
(height up to 7.2 mm, width up to 5.1 mm), 
higher than wide, conical; spire elevated, height 1,2x 
to 1.4x width; angulate periphery; umbilicus closed or 
very narrow. 
Protoconch about 180 pm wide, of 1 to 1.25 whorls, 
rounded, smooth with a thin, straight terminal lip. 
Teleoconch of up to 6.9 whorls; two first whorls 
convex, other whorls almost straight, with spiral cords 
first smooth, granular later and axial threads much 
more stronger on abapical whorls than on adapical 
whorls. Suture visible, not canaliculated. First whorl 
convex, without axial sculpture and 4 smooth cords Pi 
(i=l,2,3,4) appearing almost immediately, Pl slightly 
later and weaker; interspace between cords about 1.5 
larger than cords; ail cord brown except Pl lighter; 
axial threads very weak, hard to detect; suture 
impressed, not canaliculated. On second whorl, S2 
appearing at first midwhorl, S3 at second midwhorl, 
both quickly similar to Pi; ail cords still smooth. On 
third whorl, ail cords more or less similar in size, 
except P4 slightly stronger; distance between cords 
similar to cords; S4 may appear, partly covered by 
next whorl; weakly prosocline axial threads clearly 
visible on abapical part, especially between S3, P4 and 
S4. Shape of whorl weakly convex, almost straight 
near end. On fourth whorl, axial thread stronger 
between spiral cords; threads still weakly prosocline; 
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