L. G. Brown & B. D. Neville 
Nomenclatural notes on Amaea arabica and Cirsotrema fimbriolatum 
Therefore, Tryon and Tapparone-Canefri were 
correct in concluding that the Recent species figured 
by Kiener and Sowerby is not synonymous with the 
fossil species S. decussata Lamarck, 1804, because S. 
decussata has a different sculpture that is évident in 
the photographs of the syntypes. 
Finally, in the course of preparing this manuscript, 
we discovered there is a question regarding the 
publication date of Nyst’s Tableau, which included his 
replacement name S. arabica, as well as replacement 
names for a number of other epitoniid taxa. While it 
has generally been given as 1871, the actual 
publication date could be as late as 1874. The World 
Register of Marine Species (WoRMS) database gives 
the date of Elegantiscala arabica as 1872, though the 
source of the date is “not documented.” Nyst’s other 
names from the Tableau are cited in WoRMS as 1871. 
Nakayama (2003: 21) gives the date of Scalaria 
arabica as 1873, citing Bouiy (1913). This is 
evidently a typographical error on Nakayama’s part, 
however, as Boury (1913: 103) clearly gives 1871 as 
the date for the Tableau ; Nakayama (2003: 94) in his 
own bibliography gives the date of the Tableau as 
1871. Because the Tableau was presented at a 
December 3, 1871, meeting, we question whether it 
would hâve been published that same year. When we 
checked the Zoological Record , we discovered that 
Nyst’s Tableau appeared for the first time in the 1874 
volume, although it was listed with an 1871 date. 
While not definitive, it is an indication that the 
Tableau was published after 1871 and certainly no 
later than 1874. Buckhuys (1985) prepared a 
bibliographie note on the journals published by the 
Société Malacologique de Belgique. However, he 
began with the volume for 1872 and did not provide 
information on individual publication dates. Because 
we were unable to résolve this publication date 
question, we are following earlier authors in using an 
1871 publication date for the Tableau. 
Notwithstanding the uncertainty regarding the 
publication date, since it is clear that Nyst’s Tableau 
was published prior to Tapparone-Canefri’s 1876 
paper proposing the name C. kieneri for this Recent 
species, there is no question that the oldest available, 
and valid, name for the Recent taxon is therefore 
Nyst’s replacement name, Amaea arabica (Nyst, 
1871). 
Genus Cirsotrema Môrch, 1852: 49 
Type species: Scalaria varicosa Lamarck, 1822, by 
monotypy. 
Cirsotrema Jimbriolatum (Melvill, 1897) 
Figs 8-10, 12 
Scalaria fimbriolata Melvill, 1897: 11, pl. 6, fig. 10; 
Melvill, 1898: 2, pl. I, fig. 12. Type locality: Karachi. 
Scala ( Cirsotrema ) fimbriolata : Melvill and Standen, 
1903: 349. 
Cirsotrema kieneri: Wilson, 1993: 274, pl. 44, fig. 3; 
Weil, et. al. 1999: 128, fig. 401 (non Tapparone- 
Canefri, 1876). 
Epitonium fimbriolatum: Bosch and Bosch, 1982: 52. 
Amaea fimbriolata: Bosch, Dance, Moolenbeek, et. 
al., 1995: species no. 400. 
Amaea ( Scalina ) kieneri: Nakayama, 2003: 21, pl. 20, 
fig. 1-3 (non Tapparone-Canefri, 1876). 
Material Examined. Off Hervey Bay, Queensland, 
Australia, 61-79 m, 1 dd. Urangan, south Queensland, 
Australia, 1 dd. Broome, Western Australia, 15 m, 1 
lv. 80 Mile Beach, Western Australia, 1 Iv. 
Distribution. Gulf of Oman and Persian Gulf, east to 
Japan and Queensland, Australia. Intertidal to 79 m. 
Remarks. Cirsotrema fimbriolatum has also been 
confused with A. arabica, but differs from it markedly 
in structure, indeed belonging to a different genus. 
The original figures of Scalaria fimbriolata (Melvill, 
1897, and particularly Melvill, 1898) illustrate 
particularly well the characters of this species. 
Unfortunately the quality of the original image is not 
suitable for enlargement and reproduction here. 
While it is superficially similar to A. arabica, this 
species can be distinguished by the characteristic 
Cirsotrema costae and the base of the shell. The base 
of C. fimbriolatum is ringed by a sériés of triangular 
projections resembling teeth on a cog, whereas A. 
arabica has a simple basal ridge lacking these 
triangular projections (Fig. 1 I, 12). 
Scalaria fimbriolata is referable to the genus 
Cirsotrema because of the combination of the strong 
basal disk, spiral lirae, and the strongly crispate costae 
that consist of numerous plates that are fused together. 
This combination of teleoconch characters is présent 
in Cirsotrema varicosum (Lamarck, 1822) and a 
number of other species that hâve been referred to the 
genus Cirsotrema s.s. Species in the genus Amaea 
can hâve similar teleoconch scupture, but lack the 
crispate costae consisting of numerous fused plates. 
Acknowledgments 
We want to thank Yves Finet at the MHNG, 
Geneva, Switzerland, who furnished the photographs 
of the specimen of S. decussata (sensu Kiener) in the 
Delessert collection and authorized us to reproduce the 
figured specimens of S. decussata Lamarck in this 
manuscript. We also want to thank Kathie Way at 
NHMUK, London, England, who provided 
information on the holotype of S. arabica and 
furnished the photographs of the holotype of S. 
fimbriolata. Henry Domke photographed the 
i 11 ustrated specimen in the Bruce Neville collection. 
Alan J. Kohn ot University of Washington Biology 
provided excerpts of the catalog of Lamarck’s types. 
Emilio Garcia called our attention to the question of 
the dates of Nyst’s Tableau , and Paul Callomon at the 
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