( 313 ) 
“Combinirt treten beide Formen z. B. bei Viola auf,” u. s. w. 
In order to make this quotation more clear I add that Reinke 
found in Kerria japonica that the portion of the leaf-margin situated 
above the termination of the vascular bundle — therefore that portion 
where in many plants a so-called epithema is found — consists of 
a mucilage-secreting gland. In Prunus, as long as the leaf is young, 
each tooth of the leaf bears a dark red conical projection, which 
secretes balsam abundantly during the bud-stage. In Betala a large 
number of peltate glands (Trichomzotten), are found on the surface 
of the young leaves and internodes, while the leaf-teeth terminate in 
an egg-shaped mucilage gland. The latter only secretes in the young 
bud, whereas the peltate glands seem to function principally in the 
fully-developed leaf. In Corylus the young leaves are also covered 
over the whole of their surface with glands (Trichomzotten) which 
have the form of pedunculated buds. In addition every leaf-tooth 
also terminates in such a gland. The latter secretes in the young bud ; 
the others are only developed afterwards. 
It may be deduced from this quotation that according to Reinke 
a phenomenon, comparable to guttation, may arise in various ways, 
independently of root- or bleeding-pressure, as a result of the activity 
of glands. This activity may be due to internally situated glands 
of the leaf-margin, or to secretion of water by externally situated 
trichomes which at an earlier stage secreted mucilage (resin, balsam) 
or finally it may be the result of both internal and external glands. 
Except for the fact that the secretion of water is not always 
preceded by that of mucilage and that conversely the secretion of 
mucilage is often not followed by that of water, I have confirmed 
Reinke’s results. 
To the circumstance that in discussing his results Reinke made 
only a passing reference to secretion of water and did not refer to 
it at all in the body of his paper, nor mentioned experiments relating 
thereto, must be attributed the fact that this reference did not receive 
much attention, although his paper became widely known. 
I will now mention some observations which show that secretion 
of water in the plant can indeed be brought about by various 
As will be seen, the problem demands a more detailed and sys¬ 
tematic treatment ; I therefore intend to revert to it at a later time. 
In the first part of .this communication I will limit myself to the 
secretion of water in leaves, and return afterwards to that in 
flowers. 
