( 402 ) 
plants examined by him. In Stereospemmm hypostktum Miq.hefonnd 
them on the inside of the corolla; in other plants, viz. Clerodmdron 
Minnhassae T. et B., Heterophragma admophylhJm Seem., Nicandra 
physaloides Gartn. and Juanulloa parasitica. Ruiz et Pav. only on 
the outside and in Parmentiera cerifera Seem., Crescentia Cujete 
Linn, and Kigelia pinnata DC. both on the inside and on the outside. 
In other respects but little is known concerning these epidermal tri- 
chomes. Whether the glands which cover the outside of the calyx secrete 
water, has not been further investigated. As regards the corollar 
glands, Koorders has indeed put forward the suggestion that they 
may participate in the production of the calyx-water and that in 
the older flowerbuds the function of secreting water may pass from 
the glands of the calyx to those of the corolla, but no definite con¬ 
clusion was reached on this point; in any case the function of glands 
on the outside of the corolla had to remain hypothetical. 
Experience shows that, as was already shown above, there is not 
only a close agreement in external shape and anatomical structure 
between epidermal waterglands and mucilage- and resinglands, but 
a so between tnchomes ') which secrete water and those secreting 
nectar; hence the mere presence of glands on the corolla does not 
enable us to infer the nature of these glands. 
It is nevertheless quite possible that, in some of the plants exa¬ 
mined by Koorders, the glands on the outside of the corolla secrete 
water during the bud-stage. 
I am the more ready to assume this, because I am acquainted 
with a plant, in which, as will immediately be seen, the whole of 
the calyx water is derived from the corollar glands; for the rest this 
subject requires further investigation, because conditions may differ 
rom plant, to plant. As far as my experience extends to corollar 
to an s in plants with and without water-calyces, these glands are 
not always water-glands and in those cases, where they really are, 
i) For the sake of brevity I only refer to what has been observed by Haberlandt 
cermng t ie similarity of structure and shape of water- and nectar-secreting 
trich° meS * n connect * on w ^h the stipular nectaries of Vicia sepium and the 
obsenrar ydath £ des on the upper surface of the leaflets; further I refer to the 
cal ' * 0n l ° Koorders concerning the so-called extranuptial nectaries on the 
_ 4 _ % 9 e ia t Crescentia, Parmentiera and Stereospemmm compared with 
ese plants and to the observations of Schwendt on the 
mint, iw • , Muehlenbcckia sagittifolia and various species of Polygo- 
and b ^ amlieme and °ther species of Gossypium, Tecoma radicans 
and Ligustrum Regehanum. (in Beihefte zum bot. CentralblaU Bd. XXII Heft 3.1907.) 
water-glands ot 
extrafloral ne< 
num, Gossypi 
