(. 40 * ) 
the secretion may also take place later, when the flower has emerged 
from the bud-stage. 
This, for instance, is the case in Nicandra physaloides, to which 
further reference will be made. 
in Malva silvestris and in Sidalcea Candida, where numerous glands 
occur on the margin and on the lower surface of the petals, I have 
only observed the production of mucilage. In Hibiscus esculentus 
there is secreted, in addition to mucilage, glucose from the apical 
cell of the corollar glands; as we shall see, this case of glucose 
secretion does not stand by itself. 
I now wish to show, by means of a few examples, that the trichomes 
on the outside of the calyx are most probably always real glands, 
and that therefore in plants with water-calyces water, or mucilage, 
or both, may not only be secreted on the inside of the calyx, but also 
in other parts of the flower. Now investigation has shown, that we 
must not consider this a peculiarity distinguishing plants with water- 
calyces from other plants; we must remember that generally, in 
many plants, the same glands occur in the so-called floral region as 
on the vegetative parts and that secretory trichomes are sometimes 
found on the axis of the inflorescence, on the bracts or on the 
peduncle, on the outer or inner surface of the calyx or corolla and 
not uncommonly on the sexual organs themselves. 
I already pointed out above that on the young flowerbuds of 
Fuchsia exactly the same thin-walled, more or less club-shaped 
trichomes occur as may be found on the surface of the youngest 
leaves, and that secretion of watery mucilage also takes place on 
the surface of the flowers, when a cut plant is placed in a space 
saturated with aqueous vapour. What has here been said of Fuchsia, 
applies also to other plants ; sometimes the secretion of water, and 
sometimes that of mucilage predominates, although the water-secretion 
generally occurs in glands which originally produced mucilage. 
For the following examples 1 have selected by preference those 
which refer to plants with so-called water-calyces; as will be seen, 
they are not limited to the tropics. I first however, give an example 
°f a plant, in which nectar-secretion, mucilage-secretion and water- 
secretion can all three be observed, and in which (as I have not often 
found to be the case) mucilage and water are secreted by different 
glands. 
Finally some observations on the secretion of water by stamens 
will be added. 
