( 405 ) 
ton gland-cells. These are the “kurze, mehrzellige blasig-kopfig 
gestaltete Haare” of Nestler’s paper. 
If now, in order to convince ourselves of the secretory activity 
of these various glands we place a cut branch, bearing young flowers 
as well as young leaves, under a glass bell-jar kept moist, we can 
observe in a few hours that practically every “Sekretions Papille” 
of the peduncles, petioles, stipules and youngest leaves bears at its 
apical cell a large drop of mucilage, clear as a crystal, while the 
leaves are further covered with a watery layer of mucilage. The 
older full-grown leaves are almost dry on iheir upper surfaces, 
while their lower surfaces are covered with large drops, as already 
described by Nestler. 
From the young flowers under the bell-jar heavy drops soon hang 
down, and are replaced when they are removed by filter-paper. 
The liquid does not contain glucose, nor is it mucilaginous. .It is 
water, which is secreted by the smaller glands of elliptical shape, 
which behave from the beginning like true water-glands. Even there, 
where these glands occur on the very youngest leaves and flower- 
buds, they never form subcuticular mucilage or resinoqs products. 
The outer wall of the cells is always very thin; the cuticle is never 
seen to be raised by a substance formed inside the wall. 
Three kinds of glands can therefore be distinguished in Abutilon 
Darwinii: 
1. multicellular, thread-like trichomes, the “Sekretions-Papillen” 
of Behrens, which form the nectary at the bottom of the calyx, and 
which at the beginning*of flowering, secrete glucose with mucilage 
from the apical cell. ' 
2. glands of exactly the same structure, which secrete mucilage 
and also a little water; these are placed on the outside of the calyx 
and of the corolla, on the peduncle and petiole, on the stipules and 
on the youngest leaves. 
3. glands on the outside of the calyx and on the under surface 
of the leaves, which only secrete water. . 
In other Malvaceae the same kinds of glands are generally found, 
but with a different local distribution. 
In Malva silvestris the waterglands predominate; the leaves of 
the epicalyx, the outside of the calyx proper and the peduncles are 
thickly covered with them. 
In the moist space under the bell-jar the young flowerbuds next 
morning .have their epicalyx and calyx covered with water. The 
secreted water gradually increases to such an extent, that it literally 
drips down from the flowers. 
