IN CEYLON. 
83 
condition is where each flower has ten or more stamens, 
each with a perfect anther and occasionally a central pistil 
which ripens into fruit. 
It is obvious that a distinct correlation can be established 
in the number and position of the members of the flowers 
borne on trees of this class, and since the occurrence is so 
irregular and the stages established include the male, herma¬ 
phrodite, and female conditions, some light may be thrown 
on the evolution of sex in this genus. 
In considering the relations between the members of the 
flowers on trees of the third class we may at once eliminate 
the calyx and corolla, since in all the flowers the variation 
is only one of size, the maximum and minimum dimensions 
being in the female and male flowers respectively. 
The staminal whorl in the male flowers, whether these 
form a separate cluster or part of a monoecious or polygamous 
group, consists usually of ten stamens arranged as five pairs, 
alternating with the corolla segments. In some male flowers 
there may be only eight stamens arranged as three pairs 
and two separate individuals, and in others I have met with 
an irregular epipetalous group of stamens. In the female 
flowers, no matter where they occur, the staminal whorl is 
represented by five epipetalous staminodes disposed so as to 
be alternate with the corolla lobes. The position of the 
members of the staminal whorl in male and female flowers 
is approximately the same, and the five single staminodes of 
the female can be connected with the five groups of stamens 
usually existing in the male flowers. Since the staminode 
is really a stamen having a barren anther, we therefore see 
that the difference between the male and female flowers can 
be easily accounted for, assuming there has been abortion of 
sporogenous tissue. 
In the gynœcium the number of loculi in the ovary and 
the number and position of the ovules and seeds is the same, 
whether the pistil is part of a hermaphrodite or female 
flower. 
