410 
LOCK : STUDIES IN PLANT BREEDING 
All the above plants had coloured flowers and purple spots in the 
axils of the leaves. The four remaining plants had perfectly white 
flowers and the spots in the axils were wanting. Three of these 
plants had white testas like Satisfaction. In the fourth the colour 
was the same, but there appeared what can only be described as a 
kind of watermark, like the ghost of the maple pattern. It was 
quite distinctly visible in all the seeds of the plant. 
In the remarks which follow concerning F 3 , the same colours are 
denoted by the same letters. On white flowered plants the (invari¬ 
ably white-coated) seeds often showed the same maple watermark, 
but usually very indistinctly, and it could not always be traced in all 
the seeds of a plant even when some of them showed it. It seems 
clear that we have here a partial reappearance of the almost latent 
maple character, and in this case I should expect that on crossing 
such plants with a pure strain having pigmented testas, the full 
maple colour would make its appearance in some or all of the immediate 
offspring. It is hoped to test this conclusion by making such 
crosses during the present summer.* 
It may tend towards clearness if I here set down at length the rela¬ 
tive numbers of the different types of seeds which are to be expected 
in F 2 and F 3 on the hypothesis that the characters brown marbling 
(m) and purple spotting (p) can only appear in combination with the 
greenish or yellowish ground colour (f), but that in other respects 
Mendel’s law is followed. 
(1) Offspring of the forms showing all three colours (m. p. f.) :— 
(a) 1 constant in F 3 (m. p. f.). 
( b ) 2 giving in F 3 (m. p. f.) and (m. f.). 
(c) 2 giving inF 3 (m. p. f.) and (p. f.). 
( d) 2 giving in F 3 (m. p. f.) and (w.). 
( e ) 4 giving in F 3 (m. p. f.), (m. f.), (p. f.), and (f.). 
(/) 4 giving in F 3 (m. p. f.), (m. f.), and (w.). 
* Note added July, 1905.—In a further generation (F 4 ), grown m England 
under good conditions, the distinction between plants bearing w and those 
bearing w (m) seeds was quite clear, and the inheritance was plainly Mendelian, 
w (m) being dominant to w. Crosses of the kind mentioned have been made 
in considerable numbers, and the result will appear next year. 
