44 Wisconsin Academy of Sciences, Arts, and Letters. 
Summary. 
1. In Adiantum, the antherozoid mother cell has a dense nu¬ 
cleus containing a heavy chromatin network, and a large irregu¬ 
lar blepharoplast in the cytoplasm. Cell and nucleus grow, the 
blepharoplast elongates into a band running half way around 
the nucleus and out into the cytoplasm. Encleus and blepharo¬ 
plast move to one side" of the cell and elongate together into a 
spiral. As the elongation of the nucleus and hlepharoplast 
progress, the cytoplasm shrinks in to form a close fitting sheath. 
The cilia are attached near the anterior end of the blepharoplast 
and tend at first to extend backward about the spiral. 
2. In Aspidium falcatum, the antherozoids develop normally, 
are set free and swim about actively although they do not func¬ 
tion. 
3. The young apogamous sporophyte of Aspidium falcatum 
forms by a vegetative outgrowth from the median region of the 
prothallium. There is no vegetative fusion of nuclei at this 
stage. 
4. In the prothallium of Aspidium falcatum the nuclei have 
from sixty to sixty-five chromosomes. 
5. In the nuclei of the young sporophyte sixty to sixty-five 
chromosomes are also present. This number is maintained un¬ 
changed throughout the growth of the sporophyte as is shown 
by counting in the divisions in the young sporange. The “hap¬ 
loid sporophyte” so formed is normal in all morphological re¬ 
spects. 
6. Sixteen sporemother cells are formed as in other ferns. 
These sixteen cells fuse by pairs to form eight cells which then 
enter directly upon the reduction divisions. 
7. Eight cells are found in the spore sac in synapsis, in 
post-synapsis and throughout the divisions. 
8. A maximum of thirty-two spores is found in the mature 
spore-sac. 
