626 Wisconsin Academy of Sciences, Arts, and Letters. 
spirem, would introduce fertile sources of error in all attempts 
at determining the paired condition of sporophytic chromo¬ 
somes. 
The nucleoli of these resting nuclei vary as to size and num¬ 
ber. Commonly three nucleoli of unequal size are present, but 
often only one is to be seen and in other cases as many as five 
or six smaller ones can be made out. 
In the formation of the spindle of the last mitosis preceding 
synapsis broad strongly developed polar caps appear (Fig. 2). 
The spindle ultimately assumes the characteristic bipolar form 
of the typical mitosis. 
In the metaphases and anaphases we find typical longitudinal 
splitting and separation of the chromosomes (Fig. 3). The 
chromosomes differ strikingly in their form, number and sim¬ 
plicity from those found in the corresponding phases of the 
heterotypic division. In the telophases the reconstruction of 
the resting reticulum of the daughter nuclei is brought about 
by the development of anastomoses between the daughter chro¬ 
mosomes. During this process the chromosomes retain for a 
time their boundaries even though they have become very much 
enlarged and reticulated (Fig. 4). Finally all outlines of 
the chromosomes are lost in the uniform reticulum and the 
nucleoli appear again in apparently the same number and sizes 
as before. 
In none of the stages of this last division before reduction is 
there anything to distinguish it from a typical mitosis. There 
is nothing that can be interpreted as a preparation for the reduc¬ 
tion divisions which are to follow. 
The net knots of the resting reticulum immediately preced¬ 
ing synapsis are much more irregular in size and distribution 
than those found at the corresponding stage of the preceding 
cell generation (Figs. 5 and 6). Some of these knots; are 
large and conspicuous and others are much smaller, appearing 
as fine granules connected by finer linin strands (Fig. 6). 
They arc chromatic aggregations, irregular as to size and shape, 
connected by fine irregular strands which in staining take the 
same general color as the net knots. The strands connecting 
