638 Wisconsin Academy of Sciences, Arts, and Letters. 
As the halves of the double chromosomes of the metaphase 
are being drawn apart, the second longitudinal split appears. 
By the time the single chromosomes are fully separated from 
one another this longitudinal split is conspicuous, the resulting 
halves diverging widely from one another. They remain at¬ 
tached to one another at the end to which the spindle fibers are 
attached, giving the familiar V shaped appearance as they pass 
back to the poles (Fig. 36). The short, thick form is rer 
tained in the anaphase and not until the daughter chromosomes 
have reached the poles do they begin to elongate to form the 
slender V-shaped chromosomes characteristic of the earlier 
telophases. The apex of the Vs points toward the pole and the 
limbs radiate out from this as a center. 
As the thick-V-shaped chromosomes reach the pole the limbs 
of the Vs become considerably extended and at the same time 
becomes more slender (Fig. 32). As they elongate they be¬ 
come constricted in places which gives them a knotty appear¬ 
ance (Fig. 33). It seems probable that part of these con¬ 
strictions continue to grow deeper until the chromosome is 
almost separated in several places. By this time the chromo¬ 
somes have lost their regular contour and have become so ragged 
and uneven that it is impossible to trace their individual out¬ 
lines. At this stage one or more nucleoli become visible, indi¬ 
cating a resting condition of these daughter nuclei of the hetero¬ 
typic division (Fig. 34). In this resting condition, how¬ 
ever, it may be seen that there is no anastomosing of chromo¬ 
somes to form a reticulum. The chromosomes seem to remain 
entirely free from one another although much constricted and 
extended in regions. 
The chromosomes reappear in the prophases of the homoeo- 
typic division in the characteristic V-shape of the early telo¬ 
phases of the heterotypic division (Fig. 35). They seem 
to be formed directly by a drawing together and smoothing up 
of their outlines. I could find no traces of anything that re¬ 
sembled a spirem stage following the resting condition. It is 
impossible to determine by direct observation in Smilacina that 
these chromosomes of the homoeotypic prophases are identical 
