205 
the Embryo and Seedling in the Grammeae . 
Triticum seedlings described by us. As in our younger specimens, the 
scutellum bundles do not turn upwards in the axis before joining the stele, 
but enter the coleoptile traces at once. 
The cultivated species of Hordeuni are said by Van Tieghem to have 
two bundles in the scutellum ; the wild species to have but one. We have 
verified this in the wild species H. jubatum , L. Two interpretations of this 
P 
Text-fig. 29. Hordeum vulgare, L. Four diagrams of axis in transverse section. I. Base of 
first internode; bud in axil of coleoptile. II. Top of first node and apparent insertion of scutellum ; 
sc x , sc 2 , scutellum bundles. III. Middle of first node and real insertion of scutellum. IV. Just 
below insertion of scutellum. 
distinction are possible. The cultivated varieties may be derived from an 
ancestral form with two bundles, now extinct. We might then regard the 
dual symmetry of the scutellum as a primitive character. Or the ancestral 
scutellum may have had but one bundle as in the wild species which 
survive. In that case the doubling of the bundle in cultivated forms is 
probably adaptive, a response to the demand for more vascular tissue. The 
