421 
the Secretory Tissues of the Marattiaceae. 
Both types of secretory tissue are equally abundant in the stem, leaves, 
and roots of these Ferns; it is therefore improbable that they are produced 
in connexion with a water-storing function, especially when it is remembered 
that all the representatives of this family are usually confined to localities 
where they would not be subjected to long periods of drought. 
Whether these substances possess any biological significance in con¬ 
nexion with the attacks of animals or of parasitic Fungi it is impossible 
to say. 
Summary. 
1. Lysigenous mucilage-canals were found in every genus and species 
of Marattiaceae examined. They are usually protogenetic, but occasionally 
hysterogenetic (e. g. petiole of Kaulfussia aesculifolia ; root of Angiopteris 
evecta). 
In the petiole of Angiopteris evecta the mucilage-canals may be 
lysigenous or schizo-lysigenous. 
2. Tannin-cells are widely distributed through the sporophytic tissues 
of the Marattiaceae ; they occur either as isolated tannin-sacs or grouped 
together in regular or irregular series. 
Lysigenous tannin-ducts are formed by the solution of the septa 
between adjacent tannin-cells. 
In conclusion, I desire to express my thanks to Professor J. B. Farmer, 
F.R.S., for much helpful advice and criticism. 
Literature cited. 
1 . Bitter, G.: ‘Marattiaceae’ in Engler and Prantl, Die natUrlichen Pflanzenfamilien, i. Th., 
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2 . Brebner, G. : On the Mucilage-canals of the Marattiaceae. Journ. Linn. Soc. (London), 
Bot., vol. xxx, 1895. 
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1902. 
4 . Campbell, D. H. : The Eusporangiatae. Publication No. 140 of the Carnegie Institute. 
Washington, 1911. 
5 . -: The Structure and Affinities of Macroglossum Alidae , Copeland. Ann. of 
Bot., vol. xxviii, 1914. 
6. Charles, G. M.: The Anatomy of the Sporeling of Marattia alata. Bot. Gaz., vol. li, 1911. 
7 . De Vriese, W. H., et Harting, P. : Monographic des Marattiacees, 1853. 
8. Farmer, J. B. : The Embryogeny of Angiopteris evecta , Hoffm. Ann. of Bot., vol. vi, 1892. 
9 - and Hill, T. G. : On the Arrangement and Structure of the Vascular Strands in 
Angiopteris evecta and some other Marattiaceae. Ann. of Bot., vol. xvi, 1902. 
10 . Frank, A. B. : Beitrage zur Pflanzenphysiologie. Leipzig, 1868. 
11 . Karsten, H.: Vegetationsorgane der Palmen. Schriften d. konigl. Akad. d. Wiss., Berlin, 1847. 
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