496 Bower.—Studies in the Phytogeny of the Filicales . 
This led to correspondence with the Director of the Museum at Sarawak, 
and with only very short delay I received from him a plentiful supply 
of specimens, both dry, and in spirit. They were obtained by him on 
a collecting trip to Mount Poi, Sarawak, in April, 1913. (See J. C. Moulton, 
B.Sc., F.R.G.S., Journ. Straits Branch R. A. Soc., No. 65, 1913.) From 
these it has been possible to ascertain all the essential facts of structure, 
though naturally certain developmental details must be left aside. In par¬ 
ticular the gametophyte is still unknown. My hearty thanks are due to all 
those who helped in obtaining this material for observation, and especially 
to Mr. Moulton himself. 
The best known of the published figures of the plant is that of Hooker, 
which is quoted by Diels (E. & P., i. 4, p. 337, Fig. 175). But more 
recently a photograph has been published by Christ (Geogr. der Fame, 
p. 18, Fig. 7), which gives a good idea of the appearance of the whole plant. 
There seems, however, to be some uncertainty as to the habit of this Fern, 
which probably arises from its being rather variable. The axis is elongated, 
and the internodes between the alternate leaves of various length (Fig. 1). 
It is densely clothed with silky yellow hairs, and bears many dark brown 
roots. Partly from the angles at which the petioles come off, and partly 
from the absence of soil from some of these rhizomes, it seems probable that 
some at least of them were climbers, others creeping on the ground. This 
would accord with the earlier descriptions. For Hooker (Sp. Fil. v, p. 272) 
mentions that in Java it is found ‘on trees’. Diels (E. & P., i. 4, p. 336) 
describes it as ‘ epiphytic or terrestrial ’ ; while van Rosenburgh (Malayan 
Ferns, 1909, p. 732), who probably has had the best opportunities for 
personal observation of it in nature, describes it as e creeping or subscandent \ 
The material I have examined would accord best with this last description. 
Dichotomous branching of the rhizome has not been observed. Lateral 
branches are, however, frequent; they arise on the abaxial face of the bases 
of certain leaves, but not of all (PI. XXV, Fig. 2). The position is similar 
to that of the branches observed in Lopkosoria and Metaxya , and a similar 
position ‘ on the hinder side of the stipes of each of the erect fronds ’ has 
been ascribed to like buds in Platycerium alcicorne (Higher Cryptogamia, 
p. 252). On the other hand, in Matonia (Seward, 1 . c., pp. 174 and 187, 
Fig. 6) the branching is dichotomous, as it is also in Dipteris conjugata 
(Seward, 1 . c., p. 494); and I find the same in Dipteris Lobbiana . Other 
modes of branching have not been observed in these Ferns. These data 
are in themselves interesting in their relation to the views of Velenovsky 
and of Schoute. The matter will be considered later, when the anatomical 
relations of the parts have been described. 
The leaves are strongly dimorphic. The sterile have a firm leathery 
lamina borne on a thin wiry petiole of variable length, up to as much 
as a foot. The fertile leaves are taller, and bear a narrower lamina ; 
