586 
MR. H. M. WARD ON THE MORPHOLOGY AND THE 
supported two opposite flask-shaped branchlets : in others only one, with or without 
a pyriform body in addition. Sometimes one or the other type occurred singly and 
irregularly (fig. 4). 
The flask-shaped body is sometimes open at the apex, though I have never succeeded 
in observing anything emitted from the pore. These flask-shaped appendages recall 
to mind the peculiar bodies figured by Woronin in another group of the Pyreno- 
mycetes (Sordcirici and although no grounds exist for correlating the two phenomena 
in detail, the fact is at least worth recording that the lateral pyriform bodies in 
Meliola are capable of subserving reproduction, as will be shown hereafter. 
When the hyphee or branchlets of this fungus are looked upon from above, and a 
strong light passes through from below, one often observes a minute, circular, bright 
spot, which appears to shine through the upper wall like a very small oil-drop; 
on reversing the object, so that the lower side of the hypha comes uppermost, this 
brilliant pore-like spot appears much more evident, and is clearly due to a thinning in 
the wall of the under side of the hypha, at a spot wdrere no colouring matter is 
deposited in the cell-walls, and where the contained protoplasm is placed more nearly 
in connexion with the outside (see Plate 43, fig. 7, and Plate 44, figs. 21, 40). 
Bornet apparently refers to these bright spots when he speaks of oily globules in 
the interior of the hyphse,t though he may have been speaking of actual oil-drops 
developed in the dried specimens with which he chiefly worked. If Bornet’s remarks 
refer to the bright spots here described, the facts of their appearing only on the lower 
wall, and not being altered by alcohol, &c., remain to be explained. 
Taking all the facts into account, the view seems to recommend itself that these 
bright spots are the points of attachment of the lryphse to the epidermis; if so, they 
are to be regarded as luiustoria of a very rudimentary nature. The mycelium 
certainly is attached to the surface of the leaf, though but feebly, and it appears 
suggestive that alcohol specimens are more easily detached than fresh ones, possibly 
because the protoplasm becomes contracted and rendered brittle. No other anchoring 
bodies have been observed, and one notes that the position of these brilliant spots 
accords with that of the well-developed liaustoria in Asterina,\ a genus of fungi at 
least allied to the Meliolcis. These bright points are not always present, and in 
some cases seem to be normally absent. They are very generally formed at once on 
germination, appearing on the first short tubes put forth by the spore (fig. 40), 
a condition of things which may again be compared to what occurs in Astenna,\ 
and also in Erysiphe and allied forms. || Still another point reminding us of Astenna 
and the Erysipliece is the function of the pyriform branchlets; in some cases at 
* “Beitrage zur Morph, u. Pliys. d. Pilze,” De Bary and Woronin, ser. iii., plate 5. 
t Op. cit., p. 260, and plate 21, fig. 3. 
% See my description in Quar. Jouru. Micr. Sc., October, 1882. 
§ Bornet, op. cit., plate 28, fig. 5. 
|| De Bary, “ Beitrage zur Morph, u. Pliys. d. Pilze,” 1870, R. iii., plate 12, figs. 1 and 2. 
