278 
SVEDELIUS : LIFE-HISTORY OF 
The ovules are anatropous (fig. 7) with two integuments. 
The ovarial cavities, but not the air-chambers formed by the 
split placentae, are quite filled up with mucilage before 
fertilization and during the early stages of the embryo- 
development (fig. 6). It is a secretion from the outermost 
cell-layer of the wall of the ovarial cavities, the cells of 
which are rather large and high (PI. XXIV., A, fig. 22). The 
nucleus is always situated at the inside wall of the cell. 
The layer that secretes the mucilage just inside is rich in 
starch, contrary to the condition in the other tissues of the 
ovary. The external cells bordering on the air-chambers do 
not differ from the others in the tissue of the fruit, nor do 
they form any mucilage. 
The mucilage is deeply stained by fuchsin, methyl violet, 
methyl green, and gentian violet, but not by Congo red. 
Iodine in potassium iodide and iodine-zinc-chloride only 
cause a feeble yellowish stain. Thus it appears that the 
uci age is a pectin-mucilage, like most mucilages which 
swell m water. 
cell "JT"? that ^ occurrenc e of starch in the outside 
tton “ “ e0,!B may be Nation to the aecre- 
b “ “ ‘ be “ Ueila S e - That starch is nsed to form mncilage 
the Ciata ^ 1)6611 68peeially observed in seeds, e. g., in 
ftfuzL; ® eae the seeds produce mucilage, the formation 
coats th * be6n 8tadied Rosenberg (24). In the outer 
mncil:!L f Q SeedS ’ 6speciall y in species with abundant 
T UOn : there i8 m uch starch in the 
apnarentlr 06 - 8 cannot be U8 ®d for the embryo but 
that it is mJf? C ° nnectlon with the mucilage-formation, so 
A ^ J 6mplo y ed for the formation of mucUage. 
A product 18 thus formed called « red starch ” (cf. A. Meyer 
which It am 24), Which is opposed to be an excretion 
cells or al T™ advanced sta S e > is stored in the epidermal 
starch. « ki° Urtiier U8ed * or mncilage. Also the common 
formJi sta ^ ch » occurs in some species and is trans- 
