244 
PACIFIC SCIENCE, Vol. 1, October, 1947 
of these two species agree so nearly that I 
recommend reducing M. emarginata to a 
synonym of M. hexandra, although M. hex¬ 
andra is described as having slightly longer 
petal appendages and styles, and slightly 
shorter petals and fruit than M. emarginata. 
The somewhat variable leaves of both species 
are similar in shape and size and minute re¬ 
ticulations (see Blumea, 4: 332, 340, and 
Fig. 5, 1941). 
To supplement Dr. Lam’s original de¬ 
scription of M. emarginata, here reduced to 
a synonym of M. hexandra, I figure a leaf, 
flower, and fruit (Fig. 1). I describe the 
flower, which was not seen by him, as fol¬ 
lows: 
Commonly 3 (1 + ) flowers develop at leaf axils 
on pedicels that are green and scaly and about 
1 cm. long. The 6 sepals are in 2 rows, reddish 
brown, deltoid, reflexed in fruit. The 3 outer 
sepals are 3 mm. long, thick, smooth within, cov¬ 
ered outside with tan scales, and edged with fine 
tomentum. The 3 inner sepals are thinner, 
smaller, and narrower than the outer ones, smooth 
within, covered outside and edged with fine to¬ 
mentum. The corolla is smooth, with tube 0.7 5 ± 
mm. long. The 6 petals are thin, light brown 
tinted when dry, narrow oblong, 4 mm. long, the 
edges revolute, each petal accompanied by 2 ovate 
appendages nearly equaling it. Stamens 6; fila¬ 
ments nearly 3 mm. long, widest at base and 
tapering to narrow tip; anthers nearly 2 mm. long; 
staminodes 6, rounded, edged commonly with 2 or 
3 teeth, which may or may not have long-tailed 
tips. Ovary 1 mm. high, tomentose, 10-celled; 
disk smooth, nearly 1 mm. high; style 3.7 mm. 
long, smooth: stigma a cleft disk, slightly wider 
than the style tip. 
The discovery of this tree in Foster Gar¬ 
dens and its identification with a species of 
the Eastern Hemisphere has a further sig¬ 
nificance. Other specimens collected by H. 
M. Curran on Oahu in April, 1911, and dis¬ 
tributed to some herbaria may also have come 
from Foster Gardens. This may be true of 
a Vitex collected by Curran and described 
by Lam as V. hawaiiensis H. J. Lam ( Buiten - 
zorg Jard. Bot. Bui., Ser. Ill, 3: 39-60, 
1921). Later [idem, 5:175, 1922), Lam 
withdrew this name and stated that he be¬ 
lieved the plant to be a Mexican species, 
V. mollis Kunth. 
Fig. 1. Manilkara hexandra. A, leaf; B, flower and bud; C, part of corolla, outside: 3 petals and 
3 pairs of appendages; D, 1 petal, 2 pairs of appendages, with pistil; E, part of corolla, inside: petal, 
1 pair of appendages, 1 stamen, 2 staminodes; F, pistil, from side and in cross section; G, fruit, seed 
from side and front. 
