May 1904] 
Variability of Dictyophora 
103 
veil and the apex of the pileus is truncate. In other characters 
they are very much the same. 
In a paper in 1890, Fischer 1 concludes that the five forms 
reported in the United States should all be included under Dic¬ 
tyophora phalloidea Desvaux. This species Fischer then sub¬ 
divides into four varieties. Var. Campanulata Fischer is charac¬ 
terized by the absence of a collar surrounding the pore at the apex 
of the pileus, the flattened pseudoparenchvmatous ridges on the 
outer surface of the pileus and the flattened bands of the veil. 
Var. typica Fischer has a collar at the apex of the pileus. The 
ridges on the pileus are high, even, and are pseudoparenchyma- 
tous. They continue into the collar at the apex. The veil is 
composed of cylindrical bands. Var. duplicata (Bose) Fischer 
under which Fischer places our American forms, is characterized 
by a very broad collar at the apex of the pileus. The ridges are 
very high and toothed. The pileus and ridges have a hyphal 
structure. The veil has smaller meshes than either of the pre¬ 
ceding varieties, the openings becoming smaller toward the free 
margin where it becomes almost membranous. Var. Farlowii n. 
var. Fischer differs from the others in having a non-reticulate 
pseudoparenchymatous veil. 
In a later paper 2 Fischer adds two more varieties, making in 
all six varieties of D. Phalloidea Desvaux. These two are var. 
Guyanensis, n. var. Fischer, which has the surface of the pileus 
almost smooth, and var. Lauterbachii Fischer which has low 
ridges very irregularly placed. 
Burt 3 in 1896 combined all the forms from the United States 
having a reticulate veil under one species, D. duplicata (Bose) 
Ed. Fischer. 
Fischer in a later paper 4 raises D. Phalloidea var. Farlowii 
Fischer to specific rank. This leaves D. Phalloidea Desvaux em¬ 
bracing five varieties. 
The occurrence of this common Dictyophora in unusual num¬ 
bers in the neighborhood of Madison during the summer of '1903, 
made it possible to make some interesting observations on the 
variability of the characteristic parts of the fungus. The speci¬ 
mens were found in the region lying along the south shore of 
Lake Mendota and extending a mile west of the city. The soil 
of this regian is covered with a sod shaded at places by growths 
of trees and is at other places completely open. Along a drive, 
leading through this tract, young forest trees had been planted 
1 Loc. cit. 
2 Neue untersuchungen zur vergleichenden Entwickelungsgeschichte 
und systematik der Phalloideen. Ed. Fischer,—Denkschriften der 
Schweiz. Nat. Gesell., Bd., XXXIII. 1893. 
3 Phalloideae of the United States. Burt, E. A., Bot. Gaz. Vol. 
XXII, p. 387. 1896. 
4 Untersuchungen zur vergleichenden Entwickelungsgeschichte und 
systematik der Phalloideen. Fischer, Ed,—1890. 
