1 8 
A. S. Marsh. 
NOTES ON THE ANATOMY OF STANGERIA 
PAR A BOX A. 
By A. S. Marsh, B.A., 
Frank Smart Student, GonviUe and Cains College; Senior 
Scholar, Trinity College, Cambridge. 
[With Eleven Figures in the Text.] 
Historical. 
HE vegetative anatomy of Stangeria has been the subject, wholly 
or in part, of many previous papers. Kraus (2) described in 
some detail the histological characters of the leaf, and this account 
was amplified, with special reference to the course of the foliar 
bundles, by Nestler (4). The stem is described in a paper by 
Solms Laubach (8), but this deals chiefly with the vascular supply 
to the cone and the method of branching induced by cone-formation. 
The “mesarch” bundles of the leaves are discussed in a general 
paper on Cycad petioles by Le Goc (3), not yet published, but the 
manuscript of which the author has very kindly allowed me to read. 
The most complete description of the anatomy is given by Pavolini 
(5), but there seemed still a certain number of points to be cleared 
up, and with some of these the present paper attempts to deal. 
Material. 
The material examined was chiefly a small stem with leaves 
attached, and a leaf with the adjoining portion of cortex. The 
former came from South Africa and was given me by Professor 
Seward, the latter was kindly supplied by Mr. Lynch, Curator of 
the Cambridge Botanic Garden. 
The stem was 13-8 cm. long and 4’2 cm. in diameter at its 
thickest part. Below it tapered gradually into root and above it 
bore three leaves, the largest of which was 50 cm. from base to tip. 
Of this length, from the base to the insertion of the lowest pinnae 
was 25-8 cm. and the terminal leaflet was 1F7 cm. long. Altogether 
there were 12 pinnae arranged in approximately opposite pairs. 
The lateral pinnae measured 14 to 16 cm. in length. 
Stem Anatomy. 
The stem is subterranean and there is little development of 
mechanical tissue. The cork is very thin and the single ring of 
wood is small and feebly developed, having a diameter of only 
0’9 cm. at a point where the diameter of the stem was 3 1 cm. In 
