Codonotheca & its Relationship with Cycadofilices. i 77 
free are 1J to 2 centimeters long, and 2J to 3 millimeters wide. 
The petiole is slender and long. Regarding the manner in which 
the organ was borne it was only possible at the time of writing the 
original description to state that “Two of the fossils lie side by side 
on one of the nodules in such a way as to indicate that both were 
probably attached by long petioles to a common stem. At one side, 
and at a slightly lower level, is seen a slender striated stem, buttbe 
actual connection is not observed.” One of the new specimens now 
at hand shows several of the spore-bearing organs lying on either 
side of a central stem, those toward the top being reduced in size. 
Although the preservation is not perfect, still the relation of the 
several organs to each other and to the central stem is such as to 
leave scarcely a doubt but that they are borne by terminal divisions, 
and are attached by slender petioles to the central stem. 
The spores lie, as seen in impressions of well preserved mature 
specimens, over the inner surface of the segments from base to tip, 
and are confined to a more or less well-marked depression occupying 
from one-half to two-thirds of the width of the segments. The 
spores are large, elongate eliptical, 029 to 0-31 millimeters long, 
018 to 0*20 millimeters wide, and of a brownish colour. There is 
no grouping of the spores or other indications of the location of 
sporangia, which were doubtless more or less completely immersed 
in the tissue, the dividing walls probably disappearing at maturity. 
In connection with the original description Codonotheca was 
compared with all types of spore-bearing organ known at that time 
with which it could apparently have any possible relations. The 
type was regarded as not probably referable to any one of the better 
known classes of plants, but possible to the class of Cycadofilices, 
the fructification of which was but imperfectly known. The 
Calynimatothcca Schimperi of Stur, and ApJdcbiocarpus Stur were 
mentioned as forms possibly related. Kidston’s specimens of 
Nenropteris heterophylla were also mentioned as evidence of the 
dimorphic character of the Neuropterid group. The resemblance 
of the spores of Codonotheca in size and shape to the spores (pollen- 
grains) of Dolerophylluin was noted. The intimate association of 
Codonotheca at the Mason Creek locality with the large pinnuled 
Neuropteris, N. decipiens, led to the suggestion that Codonotheca 
may be the spore-bearing organ of this particular genus and species 
of Cycadofilices. 
The conclusions thus very tentatively stated in 1903 were later 
more definitely formulated. In a letter to Professor F, W. Oliver, 
