4 
NOTULAE ENTOMOLOGICAE 1921. 
On Halyomorpha Mayr and allied Genera. 
By E. Bergroth. 
One of the few weak points in StåTs systematic arrangement of the 
true Pentatominae of the Old World (Enum. Hem. V, p. 54—69) is his 
treatment of the allied genera Tropicorypha Mayr and Halyomorpha Mayr. 
He included in the former genus several heterogeneous elements and ba¬ 
sed the separation of the two genera practically only upon the colour and 
puncturation of the connexivum, — altogether insufficient characters. 
Since that time numerous new species of this group have been described, 
and owing to the vague limits of the two genera the true systematic place 
both of the old species and of the newcomers has been very uncertain un¬ 
til Jeannel in his work on the Pentatomidaecollected during his and Al- 
luaud’s expedition to East-Africa (Paris 1913) rendered the position of 
many forms of this group clearer by dividing it into four genera 1 ), 
to which Hymenomaga Karsch (apparently unknown to Jeannel) should 
be added, if we deem it desirable to separate it from Boerias Kirk., 
which in my opinion we cannot justly do, as it was based solely on a 
secondary sexual character (the curious form of the male membrane), 
all other characters, including the structure of the male and female 
genital segments, being exactly those of Boerias. 2 ) 
J e a n n e 1’ s generic division of this group purports a progress in our know¬ 
ledge, but it is highly to be regretted that he in dividing the old genera 
and in characterizing those accepted by himself and their species paid no 
attention whatsoever to the structure of the metasternal orificia, which 
is of prime importance in the Pentatomidae and markedly constant in the 
different genera. The orificia are of different structure in several species 
of this group, and in my opinion the old genera can not be satisfactorily 
subdivided without basing the new genera on the orificia in the first pla¬ 
ce. The structure of the orificia is not apt to be influenced by the habits 
and environments of the different species, and in agreement withBrun- 
nervonWattenwyl’s views 1 regard such characters as specially suitable 
for generic criteria. Many of the characters ascribed by Jeannel to the 
genera Halycorypha and Halyomorpha can not be regarded as of generic 
importance, the best ones being those taken from the male and female 
genital segments, although not even these are so reliable as might be de¬ 
sired. The genus Halyomorpha is characterized by Jeannel as having the 
sinuosity of the male genital segment V-shaped, but he refers the species 
reflexa Sign, and annulicornis Sign, to this genus, although the sinuosity 
in these species is U-shaped as in Halycorypha. 
9 His fifth genus, Farnya Sellout., has nothing to do with this group, as I have 
shown in Ann. Soc. Ent. Belg. 1914, p. 26. —• The West-African Cimex adspersus 
Fabr., which Stål placed in Antestia, belongs to Farnya. 
*) A new Hymenomaga (perplexa Schout.) has recently been described, founded 
only on the female. It does not seem to resemble the typical species and the aut¬ 
hor 4 does not state the reason why he refers it to Hymenomaga. From Cameroon 
1 have both sexes of an apparently- undescribed Boerias of the same group. 
