2 
Tansley and Lulham. 
du Sablon (’90) and Jeffrey (’00). When the length of the primary 
forks of the rhizome have reached about three inches, continues 
the description, weaker bundles branch off from the two big ones 
and run further out in the cortex. The uppermost of these, running 
above the axile bundles, is developed more strongly, almost equalling 
the axile ones in breadth (p. 620). The stem of a fully grown plant 
shews in the main the same distribution of bundles. The number 
of peripheral ones may rise to twelve, of which the dorsal is always 
the largest. The roots arise exclusively from the outer bundles. 
This description of the origin of the two circles of bundles in 
the young plant was first shewn by Jeffrey (’00) to be incorrect. In 
reality the two flat strands into which the original vascular cylinder 
of the stem divides are not equivalent to the two similarly shaped 
inner strands of the adult rhizome, but give rise by branching in a 
tangential direction to the outer ring of strands, while the two inner 
strands are successively nipped off dorsally from the inner face of 
the ventrally situated outer strand (’00, p. 10 and PI. 7.) 
With regard to the insertion of the leaf-trace in the adult 
plant, Hofmeister’s description is the fullest that we are aware of, 
but is partly incorrect, while his figures are inadequate for a detailed 
appreciation of the relations of the different strands. 
Stenzel (’61) corrects Hofmeister's account in one respect. He 
points out that the two circles of vascular strands are often in 
connexion by means of cross branches passing between the two plates 
of sclerenchyma from the ventral inner strand to the lateral mem¬ 
bers of the outer circle (Taf. 5, Figs. 6, 17), as well as by means of 
occasional cross branches passing through the dorsal sclerenchyma- 
tous plate from the dorsal inner to the dorsal outer vascular strand 
(Taf. 5, Fig. 8), and sometimes by similar strands passing thiough 
the ventral plate (Fig. 9). He also notes that in some weak stems 
both the two sclerenchymatous plates and the two vascular strands 
of the inner ring form a closed cylinder (Fig. 12). Stenzel’s des¬ 
cription of the origin of the leaf-trace is quite general, and his 
figures, like Hofmeister’s, fail to convey a clear and detailed picture 
of the facts. We cannot bear out his statement that before the 
leaf has separated from the stem, the foliar vascular strands are 
all thread-like (cylindrical). 
Mettenius, at the close of his well-known paper on Angiopteris 
(’63) devotes a few words to this species, and remarks that when 
the lateral bud formed on the base of the petiole develops late, the 
vascular bundles of both petiole and lateral shoot are almost 
