The Uredinece. 
i35 
by Harper and Holden. However, they have observed and figured 
the equatorial plate stage giving a characteristic and interesting 
figure (Fig. 19). All of the characteristic structural features of 
this stage in algae and fungi generally are present, polar radiations, 
chromosomes, central bodies and spindles are all sharply differentiated. 
The chromosomes are rather irregular and are never found in 
a flat plate as in the typical mitotic figure, but are always more or 
less strung out along the spindle (Figs. 18 and 19). The passage to 
to the pole is also somewhat irregular. The daughter nuclei are 
organized and cell-division follows, almost immediately. Following 
cell-division the two daughter nuclei enter the prophase of the 
second division. This stage seems to be essentially the same as 
that of the first, but the figures are so small that they are difficult 
to observe with certainty. Following the second division, cell- 
division takes place and the tetrad division is complete. In 
comparing this with the tetrad division of the spore mother-cells of 
higher plants it may also be cited that the four cells produced are 
essentially spores, as they often fall apart and give rise to the new 
mycelia, usually, however, their growth is arrested and the 
sporidia are formed. The name promycelium is therefore seen to 
be somewhat misleading. 
Variations in the Promycelium of COLEOSPORIUM 
PULSATILL/E (Str.). 
In all the forms of rusts reported so far, there seems to be a 
uniformity in the production of the promycelial cells, each cell of 
the germinating telueutospore gives rise to a four-celled structure, 
the cells of which are borne one above the other forming a structure 
in the nature of a filament. During the study of several rusts 
collected on various species of Composites and Ranunculacece growing 
in the English Garden at Munich, the observation was made that 
certain variations from this typical condition seemed to have 
occurred in a species of Coleosporium which has been identified as 
C. pulsatillce (Str.) growing on Anemone Pulsatilla. I am in no wise 
sure that this variation is expected to occur frequently in the sori 
of this rust; since I have observed it only in the case here 
mentioned, I think it is not, and must be regarded as an abnormality 
or as an occasional variation. 1 It seems to be of sufficient importance, 
however, to justify the present report. In the following paragraphs 
1 The same variation has been noted on a collection of Coleosporium 
sp. growing on Vernonia noveboracensis (Willd.) in the campus 
at Indiana University during the fall of 1906. 
